Celtica gigantea subsp. maroccana (Pau & Font Quer) F.M. Vázquez & Barkworth, 2004

Vázquez, Francisco Maria, Márquez, Francisco, García, David & Oualidi, Jalal El, 2025, A taxonomic revision of Stipeae s. l. tribe (Poaceae) in N Africa, Phytotaxa 699 (1), pp. 1-174 : 93

Celtica gigantea subsp. maroccana (Pau & Font Quer) F.M. Vázquez & Barkworth ( Figure 51 (3)).

Lemma dorsal surface: Characterised by fusiform to rectangular long cells of straight, acute to oblique apex (80–)85– 155(–210) × 7–14(–17) µm, smooth walls to 2 µm thick, linear distribution. Short cells: Silica cells of 10–14(–16) × 10–12(–13) µm, circular to ovoid, frequent, often paired with cork cells, situated between long cells, distributed irregularly, frequently solitary, density 10–12(–14) silica cells/100 µm 2; Cork cells (4–)5–7(–8) × 10–12 /µm, reniform to fusiform, sporadic, always with a silica cell. Spines and hairs: Hooks absent; Prickles not studied; Macro-hairs up to 870 µm long, circular base 22–37(–42) µm, irregularly distributed across entire surface.

Lemma apical surface: Characterised by long cells similar to those of the dorsal area. Short cells: Silica cells equal in size, morphology, density and distribution; Cork cells of equal size, morphology and distribution. Spines and hairs: Hooks absent; Prickles not studied; Macro-hairs equal in size, morphology and irregular distribution to those at dorsal part, located across entire surface except at distal part of apex. Upper apex smooth, lacking spines or hairs.

Awn base surface: Characterised by long cells of acute to oblique apex up to 530 µm in twisted, flat and angled parts. Short cells absent. Spines and hairs: Hooks absent; Prickles of oblong to obovate base 22–29(–31) × (8–)9–14 µm, up to 65 µm long, located in twisted parts, flat part smooth; frequent in angled, density 4–6 prickles/ 100 µm 2; Macro-hairs absent. Stomata paracytic of Parallel-sided subsidiary cells type, of linear distribution in flat and groove between twisted parts, density 4–6(–7) stomata/ 1cm 2.

Material studied: Morocco: Issaguen, Al-Hoceima, 1700 m.a.s.l., 20-V-2021, M . Chamboleyron, ECWP n.n.; Meknès , A 50 km de Khénifra, 33,213 06, -5,92694, 1200 m.s.a.l., 11-V-2010, Zonas altas sobres suelos graníticos acompañados de Chamaerops humilis L ., R . Ferreira, R . Gavilán, C . Pinto-Gomes, D. Sánchez.Mata , F . M . Vázquez & B . Vilches, HSS n.n.

Figures

FIGURE 51. Lemma and awn surface Celtica gigantea subsp. gigantea (1), Celtica gigantea subsp. donyanae (2) and Celtica gigantea subsp. maroccana (3). A. Lemma central dorsal surface view. B. Lemma margin dorsal surface view. C. Lemma apical surface. D. Awn surface base. lc. Long cell. sc. Silica cell. cc. Cork cell. h. Hook. p. Prickle. mh. Macro-hair. s. stomata. Bars indicate 100 µm.