Celtica gigantea subsp. gigantea (Hoffmmans. ex Link) F.M. Vázquez & Barkworth

Vázquez, Francisco Maria, Márquez, Francisco, García, David & Oualidi, Jalal El, 2025, A taxonomic revision of Stipeae s. l. tribe (Poaceae) in N Africa, Phytotaxa 699 (1), pp. 1-174 : 90

Celtica gigantea subsp. gigantea (Hoffmmans. ex Link) F.M. Vázquez & Barkworth ( Figure 51 (1)).

Lemma dorsal surface: Characterised by fusiform to rectangular long cells of acute to oblique apex (60–)100–180(– 230) × (6–)7–12(–14) µm, smooth walls to 2 µm thick, linear distribution. Short cells: Silica cells of 10–12(–14) × 10–12 µm, circular to ovoid, frequent, often in pairs with cork cells, situated between long cells, distributed irregularly, frequently solitary, density (7–)8–1(–13) silica cells/100 µm 2; Cork cells 5–8(–9) × 10–12 µm, reniform to fusiform, sporadic, always with a silica cell. Spines and hairs: Hooks absent; Prickles not studied; Macro-hairs up to 900 µm long, circular base 24–38(–44) µm, irregularly distributed across entire surface.

Lemma apical surface: Characterised by long cells similar to those of the dorsal area. Short cells: Silica cells of equal size, morphology and distribution, at lower density (6–)7–12 silica cells/ 100 µm 2; Cork cells of equal size, morphology and distribution. Spines and hairs: Hooks absent; Prickles circular base 16–25 µm and up to 50 µm long, distributed at the middle of the apex, density <8 prickles/ 100 µm 2; Macro-hairs of equal size and morphology but irregularly distributed to dorsal part, located across entire surface except at distal part of apex. Upper apex smooth, lacking spines or hairs.

Awn base surface: Characterised by long cells of acute to oblique apices up to 350 µm in twisted, flat and angled parts. Short cells absent. Spines and hairs: Hooks absent; Prickles of oblong to obovate base 18–24 × 8–12 µm, up to 35 µm long, located in the angles of twisted parts, sporadic, density <2 prickles/ 100 µm 2; Macro-hairs absent. Stomata paracytic of Parallel-sided subsidiary cells type, of linear distribution in the groove between twists, density 2–4 stomata/ 1cm 2.

Material studied: Morocco: Taza, A 55 km de Ketama a Chefchaouen, 35,001 1, -4,90094, 14-V-2010, Zonas montanas junto a Cedrus atlantica (Endl.) Carriére, R. Ferreira, R. Gavilán, C. Pinto-Gomes, D. Sánchez.Mata, F.M. Vázquez & B. Vilches, HSS n.n; Tétouan, Bab Berret, Proxidades localidad. A 20 km de Chefcaouen, 34,987 5, - 4,847 22, 1380 m.s.a.l., 14-V-2010, Bosque de Quercus pyrenaica Willd., R. Ferreira, R. Gavilán, C. Pinto-Gomes, D. Sánchez.Mata, F.M. Vázquez & B. Vilches, HSS n.n.

Figures

FIGURE 51. Lemma and awn surface Celtica gigantea subsp. gigantea (1), Celtica gigantea subsp. donyanae (2) and Celtica gigantea subsp. maroccana (3). A. Lemma central dorsal surface view. B. Lemma margin dorsal surface view. C. Lemma apical surface. D. Awn surface base. lc. Long cell. sc. Silica cell. cc. Cork cell. h. Hook. p. Prickle. mh. Macro-hair. s. stomata. Bars indicate 100 µm.