Cydistus, BOURGEOIS, 1885

Kundrata, Robin, Blank, Stephan M., Prosvirov, Alexander S., Sormova, Eliska, Gimmel, Matthew L., Vondráček, Dominik & Kramp, Katja, 2019, One less mystery in Coleoptera systematics: the position of Cydistinae (Elateriformia incertae sedis) resolved by multigene phylogenetic analysis, Zoological Journal of the Linnean Society 187, pp. 1259-1277 : 1265-1270

CYDISTUS BOURGEOIS, 1885

( FIGS 2–4)

Type species: Cydistus reitteri Bourgeois, 1885 ; by monotypy.

Diagnostic redescription: Adult male with body about 5.5–10.0 mm long; frontoclypeal region declined anteriorly; antennomeres IV–XI with rami slender, more than six times as long as wide, more than twice as long as their respective stems; labrum free, strongly transverse, fully sclerotized or partly membranous; mandible elongate, narrow, sickle-shaped, slightly widened basally, moderately curved mesally, acute apically; labial palp with three articles, palpomere I strongly transverse, palpomere II about as long as wide or slightly longer than wide, terminal palpomere elongate, fusiform; prothorax 1.0–1.2 times as long as wide; pronotal disc of variable shape; elytra shortened to various extent, together 1.4–1.7 times as long as wide, about 1.6–2.0 times as long as pronotum, more or less dehiscent; hind wing about 2.2–2.5 times as long as wide, radial cell about 4.5 times as long as wide, AP 3 + 4 distinctly forked; sternite IX 1.75–2.20 times as long as wide, apically rounded or truncate; aedeagus with phallobase longer than wide; median lobe divided

Phengodidae LeConte, 1861

Cydistinae Paulus, 1972

Cydistus Bourgeois, 1885 6 spp. Microcydistus Kundrata , 1 sp. Blank & Prosvirov

Mastinocerinae LeConte, 1881

Akamboja Roza, Quintino, Mermudes 5 spp. & Silveira, 2017

Brasilocerus Wittmer, 1963 9 spp. Cenophengus LeConte, 1881 23 spp. Cephalophrixothrix Wittmer, 1976 3 spp. Decamastinocerus Wittmer, 1988 2 spp. Distremocephalus Wittmer, 1976 11 spp. Eurymastinocerus Wittmer, 1976 8 spp. Euryognathus Wittmer, 1976 2 spp. Euryopa Gorham, 1881 8 spp. Howdenia Wittmer, 1976 10 spp. Mastinocerus Solier, 1849

subgenus Mastinocerus Solier, 1849 18 spp. subgenus Paramastinocerus Wittmer, 1976 8 spp. Mastinomorphus Wittmer, 1976 15 spp. Mastinowittmerus Zaragoza-Caballero, 1984 2 spp. Neophengus Wittmer, 1976 3 spp. Nephromma Wittmer, 1976 2 spp. Oxymastinocerus Wittmer, 1963 9 spp. Paraphrixothrix Zaragoza-Caballero, 2010 1 sp. Paraptorthodius Schaeffer, 1904 3 spp. Phrixothrix Olivier, 1909 17 spp. Pseudomastinocerus Wittmer, 1963 9 spp. Ptorthodiellus Wittmer, 1976 2 spp. Ptorthodius Gorham, 1881 3 spp. Spangleriella Wittmer, 1988 1 sp. Steneuryopa Wittmer, 1986 1 sp. Stenophrixothrix Wittmer, 1963 19 spp. Taximastinocerus Wittmer, 1963 18 spp. Phengodinae LeConte, 1861

Microphengodes Wittmer, 1976 2 spp. Phengodes Illiger, 1807

subgenus Phengodella Wittmer, 1975 20 spp. subgenus Phengodes Illiger, 1807 10 spp. Pseudophengodes Pic, 1930 26 spp. Zarhipis LeConte, 1881 3 spp. into distinct dorsal and ventral lobes.

Composition: Six described and several undescribed species. Described species: Cydistus chindaaricus Bolívar y Pieltain, 1913, C. escalerai Bolívar y Pieltain, 1913, C. nigripennis Wittmer, 1979 , C. persicus Bolívar y Pieltain, 1913, C. reitteri Bourgeois, 1885 , C. zurcheri Bourgeois, 1908 (= C. reitteri Reitter, 1908 ).

Distribution: Southern and eastern Turkey, Jordan, Israel, Iraq, Iran. Its occurrence in Syria is highly likely, but has not been confirmed.