Angulosaccus Reinhard, 1944

Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A. & Glenner, Henrik, 2020, A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala), Zoological Journal of the Linnean Society 190, pp. 632-653 : 642

Genera: Angulosaccus Reinhard, 1944 (one sp.), Boschmaia Reinhard, 1958 (one sp.), Cyphosaccus Reinhard, 1958 (four spp.), Peltogasterella (three spp.).

Remarks: The family comprises the taxa that originate at node 6 in Figure 3, and also Boschmaia and Angulosaccus . There are no molecular data for Boschmaia , but its close morphological similarity to Cyphosaccus argues for its placement here. The same argument applies to Angulosaccus . With this composition, the Peltogasterellidae contain all former peltogastrid species with colonial externae ( Reischman, 1959), and we suggest that this represents an apomorphy for the new family. Colonial externae evolved elsewhere in Rhizocephala, but not with the morphological characteristics found here. It is noteworthy that Lützen et al. (2009) found Boschmaia munidicola Reinhard, 1958 in New Zealand waters, far removed from the type locality in the Caribbean. It is therefore questionable whether more than one species exists in this deep-water genus, exactly as these authors concluded for Parthenopea Kossmann, 1874 . Colonial externae are not confined to Peltogasterellidae (see Discussion), but it still represents an apomorphy for the family at this level. Owing to the ingenious studies of Ryuzu Yanagimachi, Peltogasterella gracilis (Boschma, 1927) was the first rhizocephalan in which the presence of separate sexes and cryptic dwarf males was demonstrated. These findings entered all invertebrate and parasitological text books ( Ichikawa & Yanagimachi, 1957, 1958; Yanagimachi, 1960, 1961a, b) and inaugurated modern research on the order.

FAMILY PARTHENOPEIDAE RYBAKOV & HØEG, 2013 (MONOGENERIC)

Type species: Parthenopea subterranea Kossmann, 1874 .

Diagnosis: Morphological diagnosis as by Rybakov & Høeg (2013)