Chelotrupes momus ( Olivier, 1789 ), 2008

Dellacasa, Marco & Dellacasa, Giovanni, 2008, Revision of the genus Chelotrupes Jekel, 1866 n. stat. (Insecta, Coleoptera, Geotrupidae), Zoosystema 30 (3), pp. 629-640 : 637-639

Chelotrupes momus ( Olivier, 1789) n. comb.

( Figs 1C, F, I, L ; 2C )

Scarabaeus momus Olivier, 1789: 60 . — Illiger 1800: 152; 1803: 208.

Scarabaeus momus – Fabricius 1792: 13.

Ceratophyus momus – Sturm 1843: 113.

Ceratophyus laevipennis Mulsant & Godard, 1855: 1 . — Boucomont 1912: 22 (as synonym of momus ).

Geotrupes (Minotaurus) momus – Marseul 1863: 122. — Reitter 1883: 97.

Geotrupes (Minotaurus) laevipennis – Marseul 1863: 122. — Reitter 1883: 97.

Ceratophyus (Chelotrupes) momus – Jekel 1866: 550.

Geotrupes momus – Marseul 1867: 54.

Geotrupes laevipennis – Marseul 1867: 54.

Geotrupes andalusiacus teste Deyrolle, 1869: 10 . — Boucomont 1912: 22 (as synonym of momus ).

Geotrupes (Typhoeus) momus – Reitter 1891: 184.

Geotrupes (Typhoeus) laevipennis – Reitter 1891: 184.

Ceratophyus (Minotaurus) momus – Reitter 1893: 9.

Ceratophyus (Minotaurus) momus var. laevipennis – Reitter 1893: 9.

Ceratophyus (Typhoeus) momus – Reitter 1906: 725.

Ceratophyus (Typhoeus) momus var. laevipennis – Reitter 1906: 725.

Ceratophyus (Typhoeus) momus var. momoides – Reitter 1906: 725.

Typhoeus (Chelotrupes) momus – Boucomont 1912: 22. — Winkler 1929: 1036. — Báguena Corella 1967: 260. — Martín-Piera & López-Colón 2000: 122. Typhoeus momus – Baraud 1992: 49.

TYPE MATERIAL. — Lectotype (here designated): 1 ♂ ( MNHN), bearing the following labels: 1) blue, handwritten: “ Geotrupes momus / C. Olivier / Afr. aequin.”; 2) red, printed: “ Geotrupes momus / Olivier, 1789 / LECTOTYPUS / Des. Dellacasa 2005”.

Paralectotype: same data as lectotype, 1 ♀ ( MNHN).

TYPE LOCALITY. — “Afr. aequin.” [Africa aequinoctialis; Sierra Leone?]. Olivier (1789) and Fabricius (1792) recorded as “patria typica” of this taxon respectively “ Sierra Leone ” and “ India orientali”, and Jekel (1866) “le Portugal (Algarves), l’Andalousie et la Barbarie occidentale”. Conversely the species is known only from southwestern Iberian Peninsula.

ETYMOLOGY. — Momus is the Greek deity of mockery, faultfinding, scoff and unfair criticism.

MATERIAL EXAMINED. — Spain. Algeciras, leg. A Grouvelle, 1 ex. (MNHN). — Idem , 6.IV.1906, ex coll. Schramm, 2 ex. (MNHN). — Idem , 7.IV.1905, ex coll. Schramm, 6 ex. (MNHN). — Cadiz, 3 ex. (MNHN). — Idem , 1842, leg. Gilliani, 1 ex. (MNHN). — Cadiz, Puerto Galiz, 3-9. IV.1984, leg. J. Baraud, 11 ex. (MNHN). — Cadiz, Rota, 29.XI.1973, leg. J. Baraud, 1 ex. (MNHN). — Andalusia, El Rocio, Huelva, 14.IV.1986, leg. G. & E. Dellacasa, 1 ex. (coll. MD). — Andalusia, Facinas, 4.XII.1984, leg. G. & M. Dellacasa, 41 ex. (coll. MD). — Lag. de la Janda, IV.1918, leg. Brevil, 1 ex. (MNHN). — Prov. Sevilla, Villamanrique, 1-11.X.1960, leg. J. Baraud, 27 ex. (MNHN). — Andalusia, Tarifa, Cadiz, 3.XII.1984, leg. G. & M. Dellacasa, 2 ex. (coll. MD). — Idem , 7.XII.1984, leg. G. & M. Dellacasa, 20 ex. (coll. MD). — Cadiz, XI.1972, leg. A. J. Villarreal, 5 ex. (MCSNG -PC).

Portugal. Algarve, env. de Faro, X.1970, leg. H. Coiffait, 2 ex. (MNHN). — Algarve, Moncique, ex Musaeo Harold, 4 ex. (MNHN). — Idem , ex. coll. Bedel, 1 ex. (MNHN). — Idem , 2.IV.1967, leg. M. Coiffait, 1 ex. (MNHN). — Idem, Foia , 900 m, 20.IV.1982; leg. J. Baraud, 5 ex. (MNHN). — Algarve, Sta. Barbara de N., IV.1963, leg. M. Coiffait, 8 ex. (MNHN). — Idem, XII.1963 , 2 ex. (MNHN). — Algarve, Tavira, 14-25.II.1965, leg. T. Branco (excr. humain), 5 ex. (MNHN). — Ribatejo, 16 km Sud Coruche, 3-9.IV.1984, leg. J. Baraud, 9 ex. (MNHN). — Ribatejo, 6 km N de Lavre, 29.III.1984, leg. J. Baraud, 2 ex. (MNHN). — Ribatejo, Monte da Barca, 2.IV.1984, leg. J. Baraud, 2 ex. (MNHN). — Ribatejo, Muge, 27-28.III.1984, leg. J. Baraud, 3 ex. (MNHN). — Idem , 6.IV.1981, leg. J. Baraud, 1 ex. (MNHN). — Idem , 7.IV.1986, Leg. G. & E. Dellacasa, 1 ex. (coll. MD).

DISTRIBUTION. — Known from southern Spain and Portugal (Ribatejo and Algarve). We have examined specimens preserved in the Paris Museum coming from localities that need to be confirmed: Maroc, ex coll. Boucomont, 3 ex.; Algérie, ex coll. Ernest André, 1914, 2 ex. The record from Ibiza, Balearic Islands ( Crovetti 1970) needs to be confirmed too.

BIONOMICS. — Flightless telocoprid species utilizing several kinds of dung (sheep, horse, cow), particularly frequent in dung of rabbits (Sanchez-Piñero & Avila 1991), rarely in dung of wild boar ( Hidalgo et al. 1998) and of lynx (Martín-Piera & López-Colón 2000), also in human excrement. Present all year round and distributed from sea level to high hills.

DESCRIPTION

Length 10.0-21.0 mm, moderately stout, strongly convex, shiny, glabrous. Black; antennae and legs blackish. Head with epistoma narrowly rhomboidal, glabrous in both sexes, with median longitudinal gibbosity moderately raised and more or less coarsely punctate; clypeus subtriangular, rounded at anterior angle, thinly margined, the edge sparsely ciliate and distinctly upturned distally; genal sutures cariniform, strongly raised; genae widely rounded; front smooth on disc, moderately punctate laterally. Pronotum strongly transverse, moderately convex, with distinct antero-median rounded foveola on sides; latter feebly arcuate, thinly margined, the edge sparsely ciliate; hind angles widely rounded; base thinly margined, not sinuate.Scutellum subregularly triangular, almost convex. Elytra distinctly longer than wide, strongly convex, narrowly flattened before the epipleural carina; the latter glabrous; striae almost obsolete with nearly imperceptible punctures; intervals flat, smooth. Fore tibiae apical spur feebly sinuate. First segment of hind tarsi as long as following two combined.

Male: epistoma with median longitudinal gibbosity feeble, finely and very superficially punctate on disc, punctation coarser distally; pronotum superficially punctate on sides toward the anterior angles, anteriorly with three hornlike apophyses; the lateral ones straight, anteriorly projecting from above the anterior angles, subparallel at outer margin and with distinctly raised longitudinal carina subtuberculiform at middle and from this abruptly recumbent toward the inner margin that is sinuately widened inwardly, the groove distinctly rugulose; the central apophysis shortened, regularly acuminate, upturned; aedeagus illustrated on Figure 2C .

Female: epistoma with median longitudinal gibbosity distinctly tuberculiform at top, strongly, rugosely, confusedly punctate throughout; pronotum sparsely and superficially punctate on disc, punctation irregular and more dense on sides, with antero-lateral angles dentiform, feebly projecting anteriorly and distinctly outwardly curved; at each side foveolate and with an outer tooth sharp and distinctly raised; with short strong antero-median transverse carina.

Figures

FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb.

FIG. 2. — Aedeagus in dorsal view: A, Chelotrupes hiostius (Gené, 1836) n. comb., Costa Verde, Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; B, C. matutinalis (Baudi, 1870) n. comb.,Costa Verde,Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; C, C. momus (Olivier, 1789) n. comb., Andalusia, Cadiz, Tarifa. Scale bar: 1 mm.