Coleostephus Cass.

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 174

80. Coleostephus Cass. 1

Annuals. Leaves simple, alternate. Capitula solitary or 2-5 on branches. Involucral bracts in 2-3 rows, brownish with a narrow, scarious apical border. Receptacle plano-convex, without scales. Ligulate florets female, fertile or sterile. Inner florets hermaphrodite, tubular-campanulate, actinomorphic; corolla-tube strongly compressed at base and slightly winged. Achenes all similar, cylindrical, with 8-10 more or less projecting white ribs and a basal callus. Epicarpic mucilaginous cells on some ribs and vallecular secretory canals present. Pappus prominent, oblique, sheathing or a corona, as long as or shorter than achene, sometimes absent.

Leaves±regularly toothed; corolla-teeth of tubular florets subacute; pappus an auricle subequalling achene, sheathing 1. myconis Leaves irregularly toothed; corolla-teeth of tubular florets truncate; pappus a corona or absent 2. clausonis