Knautia macedonica, Griseb.

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 66

32. K. macedonica Griseb. , Spicil. FI. Rumel. 2: 178 (1846)

( K. atrorubens Janka ex Brandža ).

Like 29 but leaves evenly distributed along the stem, the basal usually withered at anthesis, undivided, the cauline with ovate, crenate-serrate terminal lobe; capitula 1-5-3 cm in diameter; corolla usually dark red, sometimes lilac or pink. 2и = 20. Scrub and open woods. • C. part of Balkan peninsula, S.E. Romania. Al Bu?Gr Ju Rm.

Biennial variants with wide-based cauline leaves, from S.E. Romania, have been named K. tulceanensis E. I. Nyárády , Bul. Grad. Bot. Univ. Cluj 19: 82 (1939). Populations are often polymorphic in the colour of the corolla, possibly because of hybridization, and approach 29 and 31.