Masuria

Assing, Volker, 2006, New species and records of Masuria Cameron, with a key to species, Beiträge Zur Entomologie = Contributions to Entomology 56, pp. 141-154 : 150-153

Key to the species of Masuria

In the following key, references to illustrations in the literature are abbreviated as follows: A98 = A s s in g (1998); A04 = A s s in g (2004); P89 = P a c e (1989); P98 = P a c e (1998). The distinguishing characters of M. fe r r u g in e a, M. ru b id a, M. ru fescem, M. b esu ch eti, and M. tashigaonensis, of which no material was seen, are based on the descriptions and keyprovided by Pace (1989).

1. Elytra very short, at most 0.7 times as long as pronotum; hind wings reduced. Lateral margins of pronotum distinctly dilated, abruptly convex to distinctly angled in anterior half. Posterior angles of pronotum well-marked. Abdomen at least as wide as elytra or nearly so; posterior margin of tergite VII without palisade fringe. Subgenus Oncosomechusa 2

Elytra longer, distinctly more than 0.7 times as long, usually approximately as long as pronotum; hind wings present. Lateral margins of pronotum more or less evenly convex in dorsal view, sometimes with maximal width in anterior half. Abdomen distinctly narrower than elytra; posterior margin of tergite VII with palisade fringe. Subgenus Masuria 7

2. Species from the Himalaya (Nepal and northern India).......................................................... 3

Species from China........................................................................................................................5

3. Coloration of body yellowish red to reddish. Aedeagus and spermatheca as in Figs. P89: 44-47. Central Nepal.................................................................................. M. m artensi (Pace)

Coloration of body darker brown................................................................................................ 4

4. Eyes larger, more than half the length of postocular region in dorsal view. Pronotum less transverse (Fig. P89: 39). Spermatheca as in Fig. P89: 40 (male unknown). Northern India: Darjeeling district........................................................................................ M. b esu cheti (Pace)

Eyes very small, less than half the length of postocular region in dorsal view. Pronotum more transverse (Fig. P89: 42). Spermatheca as in Fig. P89: 41 (male unknown). Nepal: Khandbari district.................................................................................. M. tashigaonensis Pace

5. Elytra approximately 0.7 times as long as pronotum. Pronotum distinctly dilated in anterior half (Fig. A04: 19). Puncturation of forebody distinct and well-defined. Eyes longer than postocular region in dorsal view (Fig. A04: 19). Antennomere IV distinctly (approximately 1.5 x) oblong. Legs, especially metatarsus, long and slender. Abdomen without microsculpture. Aedeagus as in Fig. A04: 20. China: N-Yunnan....................... M. yunnanica Assing

Elytra shorter, less than 0.6 times as long as pronotum. Pronotum with evenly convex lateral margins in dorsal view. Puncturation of forebody shallow. Eyes distinctly shorter than postocular region in dorsal view. Antennomere IV approximately as long as wide. Legs shorter. Abdomen with microsculpture...................................................................................... 6

6. Elytra approximately 0.55 times as long as pronotum. Antennae distinctly longer and more slender; antennomeres V-VII not or only indistinctly transverse. <?: tergite VII with broadly and distinctly concave (!) posterior margin. Aedeagus as in Figs. P98: 2-3. Spermatheca as in Fig. P98: 4. Gansu: Xinlong Shan.......................................................... M. chinensis Pace

Elytra approximately 0.50 times as long as prontoum (Fig. 34). Antennae shorter; antennomeres V-X distinctly transverse (Fig. 36). Spermatheca as in Fig. 38 (d unknown). N- Yunnan: surroundings of Zhongdian.................................................... M. b revip en n is sp. n.

7. Species from northern India with rufous to light brown pronotum and elytra................... 8

Species unknown from northern India and/or with blackish pronotum.......................... 10

8. Larger species. Elytra approximately as long as pronotum or nearly so................................. 9

Smaller species. Elytra somewhat shorter than pronotum (Fig. P89: 30). Aedeagus as in Figs. P89: 31-32. Chakrata district, Mussoori.............................. M. fe r r u g in e a Cameron

Eyes smaller, shorter than postocular region in dorsal view. Pronotum less transverse; lateral margins with long setae (Fig. P89: 27). Aedeagus as in Figs. P89: 28-29. Ghum district. ...................................................................................................................... M. ru bida C a m e r o n

Eyes larger, as long as postocular region in dorsal view. Pronotum more transverse; lateral margins without long setae (Fig. P89: 24). Aedeagus as in Figs. P89: 25-26. Ghum district........................................................................................................... M. ru fescen s C a m e r o n

Pronotum relatively large, at least 1.4 times as large as head, and with conspicuously coarse, rugose, and confluent puncturation; surface almost completely matt. Coloration of body predominantly black. Eastern Nepal: Rolwaling Himal.................... M . sculpticollis sp. n.

Pronotum in most species distinctly less than 1.4 times as large as head; puncturation not conspicuously rugose, finer, and in most species well-defined and with glossy interstices. Species with a conspicuously coarsely and densely punctate pronotum are ofbrown to dark brown coloration......................................................................................................................... 11

Species from China. Flagelloid structure in internal sac of aedeagus long and with small base (Fig. A04: 14-15). Yunnan: Dali...................................................... M. daliensis A s s in g

Species from the Himalaya. Flagelloid structure in internal sac of aedeagus usually shorter and with larger and often more or less anchor-shaped base. (For a reliable identification of the following species, an examination of the aedeagus is usually essential.) ................... 12

Pronotum with coarse and very dense puncturation; interstices very narrow, microsculptured, and either with reduced or almost completely without shine............................................. 13

Pronotum with more or less dense, but not coarse puncturation; interstices usually without microsculpture and with some shine....................................................................................... 15

Eyes longer than postocular region in dorsal view. Aedeagus relatively small and with relatively weakly pronounced crista apicalis (Figs. P89: 10-12, A98: 4h). Widespread species: Nepal, northern India........................................................................... M. p lu m b ea C a m e r o n

Eyes shorter than postocular region in dorsal view. ...............................................................14

Aedeagus much larger and with relatively short ventral process; base of flagelloid structure small (Figs. A98: 4a-b). Nepal: Khandbari district..................... M. ru gosepu nctata A s s in g

Aedeagus small and with long ventral process; base offlagelloid structure larger (Figs. P89:

18-19). Northern India: Ghum district................................................. M. p a rva C a m e r o n

Pronotum with shallow, dense, and partly ill-defined puncturation................................. 16

Pronotum with distinct, more or less dense, and well-defined puncturation.................. 19

Preapical antennomeres (IX-X) distinctly transverse, approximately 1.5 times as wide as long (Fig. 17). Flagelloid structure in internal sac of aedeagus with very large base (Figs. 21-23). Nepal: Rolwaling Himal..................................................................... M. follita sp. n.

Preapical antennomeres (IX-X) weakly transverse, less than 1.5 times as wide as long. Flagelloid structure in internal sac of aedeagus with smaller base........................................... 17

Median lobe of aedeagus smaller and in lateral view with straight ventral process; ventral process without pronounced lateral folds (Figs. A04: 6-7). Nepal: Annapurna, Dhaulagiri............................................................................................................... M. a n n a p u rn a e A s s in g

Median lobe of aedeagus distinctly larger and in lateral view with slightly bent ventral process; ventral process with lateral folds............................................................................... 18

18. Median lobe of aedeagus in lateral view dented between base of ventral process and crista apicalis (Figs. P89: 22-23). Nepal: Bagmati province ............................... M. spectata P a c e

Median lobe of aedeagus in lateral view not dented between base of ventral process and crista apicalis (Figs. 7-11). Nepal: Rolwaling Hiraal............................... M. kleebergi sp. n.

19. Pronotum with moderately dense puncturation; width ofinterstices on average at least half the diameter ofpunctures and distinctly glossy. Aedeagus as in Figs. P89: 2-3. Central and e a ste rn Nepal........................................................................................................... M. lo eb li P a c e

Pronotum with very dense puncturation; interstices narrower than half the diameter of punctures and with subdued shine............................................................................................ 20

20. Lateral margins ofpronotum each with approximately 5 long setae, which are approximately as long as antennomere III. Aedeagus very large (Figs. P89: 14-15). Nepal: Khandbari district........................................................................................................................ M. kali P a c e

Lateral margins of pronotum each with fewer (if any) and shorter setae. Aedeagus smaller...................................................................................................................................................... 21

21. Whole body, including abdomen, very densely punctate, almost matt. Abdomen (including tergite VII) with relatively coarse and dense puncturation, interstices distinctly narrower than punctures. Pronotum relatively large, approximately 1.2 times as wide as long (Fig. P89: 34). Lateral margins ofpronotum in posterior halfnot sinuate in dorsal view. Aedeagus as in Figs. P89: 35-36. Central and eastern Nepal.................................... M. sm etanai Pa c e

Body less densely punctate and with shiny interstices. Abdominal tergite VII with much finer and sparser puncturation, interstices distinctly wider than diameter of punctures. .. 22

22. Antennae shorter, antennomere X more than 1.5 times as wide as long. Legs shorter; metatarsomere I barely as long as the combined length of II and III; III approximately twice as long as wide. Coloration darker, femora blackish. Aedeagus as in Figs. P89: 6-7, A98: 3h. Widespread species: Nepal, northern India.......................................... M. p icip es C a m e r o n

Antennae longer, antennomere X less than 1.5 times as wide as long. Legs longer; metatarsomere I at least as long as - usually longer than - the combined length of II and III; III clearly more than twice as long as wide. Coloration paler, femora brown.........................23

23. Pronotum more transverse, approximately 1.25 times as wide as long. Metatarsus shorter and with less elongated metatarsomere I. Aedeagus with shorter ventral process; flagelloid structure in internal sac with more slender and anchor-shaped base (Figs. A98: 5a-b). Nepal: Rasuwa district..................................................................................... M. a n co rifo rm is A s s in g

Pronotum less transverse, approximately 1.15 times as wide as long. Metatarsus long and with elongated metatarsomere I. Aedeagus with longer ventral process; flagelloid structure in internal sac with more massive base (Figs. A98: 3a-b). Nepal: Khandbari district.......... ................................................................................................................... M. lo n g ico m is A s s in g