Asioclinocera, Saigusa & Sinclair Introduction, 2022

Saigusa, Toyohei & Sinclair Introduction, Bradley J., 2022, A new genus of Clinocerinae (Diptera: Empididae) from Taiwan and Yunnan (China), Bonn zoological Bulletin 71 (2), pp. 177-183 : 177-178

publication ID	https://doi.org/10.20363/BZB-2022.71.2.177
publication LSID	lsid:zoobank.org:pub:68F9C120-14D4-4562-B3FB-28917927E795
persistent identifier	https://treatment.plazi.org/id/8774C650-133A-8D78-5C7E-D146EDFA021D
treatment provided by	Felipe
scientific name	Asioclinocera
status	gen. nov.

Treatment

Asioclinocera View in CoL gen. nov.

urn:lsid:zoobank.org:act:DDB82413-42C5-4792-A5EF-26F9AD8AF9BF

Figs 1–10 View Figs 1–3 View Figs 4–5 View Figs 6–7 View Figs 8–9 View Fig

Type-species

A. yunnanica gen. et sp. nov.

Diagnosis. This new genus is distinguished from all other genera of Clinocerinae by the following combination of characters: elongate, filamentous arista-like stylus, face with deep cleft and ridge, labrum convex, radial fork elongate with vein R 4+5 branching near apex of cell dm and cell r 4 about one-third length of wing, apical fifth of fore femur with spur-like anteroventral seta, male tergum 8 greatly narrowed medially and broadly plate-like laterally, and phallus broadly attached to hypandrium with large ejaculatory apodeme.

Etymology. The generic name is derived from Asia, referring to its distribution, and Clinocera Meigen, 1803 , a related genus. The gender is feminine.

Description. Eye with dense ommatrichia. Arista-like stylus very slender and long, about as long as head height ( Fig. 2 View Figs 1–3 ). Face with median ridge on lower half. Gena divided by frontoclypeal suture extending from palpus to lower margin of eye. Mouthparts with labrum convex ( Fig. 2 View Figs 1–3 ); labellum sucker-like, without pseudotracheae. Fore femur with preapical anteroventral spine-like seta, slightly shorter than width of femur ( Fig. 2 View Figs 1–3 ); without subapical anterior comb, only cluster of short stiff setae; anteroventral and posteroventral rows of slender short setae ( Fig. 5 View Figs 4–5 ). Fore tibia with anteroventral row of short stout setae ( Fig. 5 View Figs 4–5 ). Tarsomere 5 without apical prolongation. Hind tibia with distinct anteroventral seta on apical third ( Fig. 1 View Figs 1–3 ). Scutum with long dorsocentral setae ( Fig. 5 View Figs 4–5 ); without acrostichal setae; notopleuron with weak anterior seta and strong posterior seta; scutellar setae very closely approximated at apex (width of anterior ocellus); laterotergite with setae; dorsal mesepimeral pocket present. Wing with basal costal seta; without pterostigma ( Fig. 3 View Figs 1–3 ); Sc complete; costa circumambient with strong, erect setae beyond R 1; R 1 ends before middle of wing; elongate cell dm; radial fork very long, arising near apex of cell dm; R 4 slightly arched to straight and divergent from R 5; cell r 4 about one-third length of wing; cell cua narrow, with truncate apical margin. Female abdomen telescopic, apparently unmodified, with cerci of A. yunnanica (unknown for formosana ) short, shiny, sharply pointed (not dissected). Male terminalia ( Figs 6–7 View Figs 6–7 ) with setae of cercal plate not confined to small region ( Fig. 8 View Figs 8–9 ); phallus without distiphallus; base of phallus and hypandrium expanded; male tergum 8 narrow with inflated lat- eral margin bearing many setae ( Fig. 9 View Figs 8–9 ).

Distribution. This new genus is currently known from Taiwan and the Chinese province of Yunnan ( Fig. 10 View Fig ).

Remarks. This new genus of Clinocerinae appears to be most similar to Rhyacodromia on the basis of the convex-shaped labrum and spine-like setae on the fore femur. The convex labrum is a rather unique character within Clinocerinae and previously considered a synapomorphy of the clade Rhyacodromia + Trichoclinocera Collin, 1941 ( Sinclair 1995, character 25). However, the male terminalia distinctly differ between Rhyacodromia and Asioclinocera gen. nov., specifically the connection of phallobase and hypandrium is unique in the former genus (anterolateral corner of hypandrium has unique one-point articulation with the anteroventral corner of epandrium and posterolateral corner of sternum 7 ( Saigusa 1986: fig. 9; Sinclair 1995: fig. 15)), whereas the connection in Asioclinocera gen. nov. is similar to Clinocera and Wiedemannia Zetterstedt, 1838 ; the surstylus is widely connected with the distal margin of the epandrium in Rhyacodromia compared to the narrow connection with the epandrium in Asioclinocera gen. nov.; and tergum 7 bears a denticulate posterior swelling in Rhyacodromia and is unmodified in Asioclinocera gen. nov. The new genus is further distinguished in the following identifi- cation key.