Holcostethus Fieber, 1860

Belousova, E. N., 2007, Revision of the shield-bug genera Holcostethus Fieber and Peribalus Mulsant et Rey (Heteroptera, Pentatomidae) of the Palaearctic Region, Entomological Review 87 (6), pp. 701-739 : 729-730

Subgenus Holcostethus Fieber, 1860

Diagnosis. Jugae as long as clypeus. Pygofer small, with widely rounded apical angles and shallowly emarginate apical margin ( Fig. 64 ). Conjunctive proc-esses of aedeagus with sclerotized apices ( Figs. 39, 40 ). Paramere in form of overturn “L,” flat; hypophy-sis with tooth and with more or less strongly devel-oped uneven area of inner surface. Spermatheca with spherical bulb bearing long processes projecting be-yond proximal flange and with long sclerotized duct slightly widened closer to base. Laterotergite IX with widened and subrectangular apex; sternite IX rather long, with straight base and pointed anterior angles.

Composition. The subgenus includes three Mediterranean species: H. albipes , H. sphacelatus , and H. evae .

Figures

Figs. 63–73. (63–67) Pygofer [(63) Paraholcostethus breviceps (Horv.), (64) Holcostethus albipes (F.), (65) Holcostethus evae Ribes, (66) Peribalus classeyi Hob., (67) Holcostethus (Enigmocoris) fissiceps Horv.]; (68–73) parameres [(68) Paraholcostethus breviceps (Horv.), (69) Himalayacoris pilosus sp. n., (70) Peribalus classeyi Hob., (71) Peribalus imtiazi Abbasi, (72) Peribalus lodosi Ahmad, (73) Peribalus sinuatus Ahmad)].

Figs. 35–40. Aedeagus, front and lateral view: (35, 36) Himalayacoris pilosus sp. n., (37) Peribalus hoberlandti Lodos et Önder., (38) Paraholcostethus breviceps (Horv.), (39, 40) Holcostethus sphacelatus (F.).