Holcostethus albipes (Fabricius, 1781)

Belousova, E. N., 2007, Revision of the shield-bug genera Holcostethus Fieber and Peribalus Mulsant et Rey (Heteroptera, Pentatomidae) of the Palaearctic Region, Entomological Review 87 (6), pp. 701-739 : 731

Holcostethus albipes (Fabricius, 1781)

Cimex albipes Fabricius, 1781: 345 .

Holcostethus congener Fieber, 1861: 334 (syn. Stål, 1868: 29).

Peribalus albipes : Puton, 1882: 58.

Holcostethus albipes : Tamanini, 1981: 133.

Diagnosis. H. sphacelatus differs from the closely related species in the smaller body, clearly concave sides of the pronotum bordered with a pale costa, and larger spermathecal bulb bearing shorter processes occasionally furcate at the apices; hypophysis of the paramere forms no constriction in the middle part, bears a tooth at the base ( Fig. 50 ).

Description. Body pale yellow to brownish, with coarse black punctation (distance between punctures exceeding, or equal to diameter of puncture). Underside of body pale, with rather sparse punctation. Abdominal segments with black spots in anterior and posterior corners of tergites. Antennal segments dark, with pale bases. Body 7–8.5 (7.72) mm long, rather robust, b. lw 1.50–1.65 (1.57).

Head distinctly narrowed anteriorly, h. wl 1.12–1.21 (1.17), medium-sized, l. bh 3.95–4.36 (4.14). Jugae with narrowly rounded apices, slightly emarginate before eyes, as long as clypeus, not covering latter ( Fig. 94 ). Eye large, ds 1.46–1.6 (1.5). Antenna long, l. ba 1.77–2.11 (1.93).

Pronotum with concave sides bordered with pale costa, p. wl 2.21–2.34 (2.25). Lateral angles widely rounded, slightly projecting beyond base of elytra. Scutellum of subequal length and width, s. lw 0.96– 1.05 (1.01).

Male genitalia. Hypophysis of paramere without constriction in middle part, with small tooth at base ( Fig. 50 ).

Female genitalia. Spermatheca with spherical bulb (r. bb 10.1) bearing processes occasionally branching at apices ( Fig. 13 ). Laterotergite IX moderately widened at apex ( Fig. 101 ).

Distribution. Mediterranean basin: Spain, southern France, including Corsica, Italy, Malta, the Balkan Peninsula, Algeria, Morocco, Syria.

Material examined: 3 Ô, 3 ♀, Positano , Campania, Italia, VII.1929 (El. Miram) ; 1 Ô, 2 ♀, Trieste, V– VIII.1929 (Passauro) ; 1 ♀, Asuni, Sardinia centr., V.1910 (A. Krausse) .

Figures

Figs. 50–62. Parameres: (50–53) Holcostethus Fieber [(50) H. albipes (F.), (51) H. sphacelatus (F.), (52) H. evae Ribes, (53) H. (E.) fissiceps Horv.], (54–62) Peribalus Mulsant et Rey [(54) P. strictus capitatus (Jak.), (55) P. strictus vernalis (Wolff), (56) P. strictus strictus (F.), (57) P. congenitus Putsh., (58) P. hoberlandti Lodos et Önder, (59) P. nitidus (Kir.), (60) P. inclusus (Dohrn), (61) P. manifestus Kir., (62) P. tianshanicus sp. n.].

Figs. 94–99. Head: (94, 95) Holcostethus Fieber [(94) H. albipes (F.), (95) H. sphacelatus (F.)]; (96) Himalayacoris pilosus sp. n.; (97) Paraholcostethus breviceps (Horv.); (98, 99) Peribalus Mulsant et Rey [(98) P. inclusus, (99) P. przewalskii sp. n.].

Figs. 12–17. Spermatheca: (12) Holcostethus sphacelatus (F.), (13) Holcostethus albipes (F.), (14) Peribalus urakensis Ahmad et al., (15) Peribalus manifestus Kir., (16) Paraholcostethus breviceps (Horv.), (17) Peribalus hoberlandti Lodos et Önder.

Figs. 100–105. Genital segments of female: (100, 101) Holcostethus Fieber [(100) H. sphacelatus (F.), (101) H. albipes (F.)]; (102–105) Peribalus Mulsant et Rey [(102) P. strictus vernalis (Wolff), (103) P. inclusus (Dohrn), (104) P. tianshanicus sp. n., (105) P. hoberlandti Lodos et Önder].