Elsholtzia winitiana Craib

Bongcheewin, B., Chantaranothai, P. & Paton, A., 2015, Elsholtzia (Lamiaceae) in Thailand, Blumea 59 (3), pp. 209-214 : 213

8. Elsholtzia winitiana Craib

Elsholtzia winitiana Craib (1918) 368; Doan (1936) 980; Murata (1971) 497; (1976) 184; Press (1982) 70; X.W. Li & Hedge (1994) 249; Phuong (1995) 35; A.L. Budantzev (1999) 21; Phuong (2000) 235. — Type: Kerr 1607B (holotype K (http://specimens.kew.org/ herbarium/K000929591, last acc. 5 Mar. 2015); isotypes E, 2 sheets K (http://specimens.kew.org/herbarium/ K000192605 & http://specimens.kew.org/ herbarium/K000929592, last acc. 5 Mar. 2015), Thailand, Chiang Mai, Doi Sutep, mixed jungle, 660 m, 3 Dec. 1911.

Shrubby perennial herb up to 1.5 m tall. Stems erect, pale green, not shiny, densely pubescent. Leaves elliptic-ovate or elliptic, 4–12 by 1.5–3 cm, base cuneate, margin shallowly serrate; upper surface densely pubescent on venation; lower surface densely pubescent with translucent pale yellow dots; petioles 10–30 mm long. Inflorescence terminal and axillary, much branched, compact, adjacent verticils overlapping, or lax, verticils from adjacent nodes not overlapping, 5–8 mm apart, cylindrical, 4–9 cm long; bracts narrowly lanceolate, 0.5–1 by 2.5–3 mm, pubescent on both sides; bracteoles linear, 2 mm long. Calyx campanulate, 1.8–2.3 mm long at anthesis, densely pubescent with translucent brown dots outside, glabrous inside; tube ventricose, 1–1.5 mm long; lobes unequal, triangular or triangular-ovate. Corolla white, 1.5–2 mm long; tube glabrescent outside, glabrous with an annulus inside; lateral lobes ovate, apex truncate; median lobe broader than lateral lobes; lower lip elliptic-ovate, hispidulous with translucent brown dots outside, glabrescent inside. Stamens 4, slightly exserted, glabrous. Style slightly exserted. Nutlets 4, oblong, 0.8 mm long.

Distribution — China, Myanmar, Thailand (N: Chiang Rai, Chiang Mai, Nan, Uttaradit), Vietnam.

Ecology — Open places in hill evergreen, deciduous, pine or dipterocarp forests, 600–2000 m altitude. Flowering and fruiting: October to March.

Specimens examined. Bongcheewin 234, 480, 595, 934; Chayamarit et al. 2360, 3251; Chermsirivathana 317, 749; Chueachom 497; Hansen et al. 10789; Jatupol 08­456; Kerr 1607, 1607B; Koyama & Phengklai T­40034; Koyama et al. T­31975, T­32237, T­32279, T­32281; Maxwell 97­1492, 98­ 1426, 04­777; Nanakorn et al.10302; Paisooksantivatana 590­81; Phengklai et al. 6271; Ploenchit 971; Sadakorn 636; Sangkhachand 126; Sankamethawee 381; Santisuk 6648; Smitinand & Alsterlund 6729; Smitinand et al. 10282; SØrensen et al. 1297, 2746; S.P. 45; Suddee et al. 2910; Tagawa & Yamada T­79.

Note — Elsholtzia winitiana is morphologically similar to E. pubescens Benth. , by having a compact inflorescence, membranaceous leaf texture and shrubby habit. However, E. winitiana has a smaller corolla, 1.5–2 mm long and is found from China and Indo-China, whereas E. pubescens has a larger corolla, c. 4 mm long and found in Java, Lesser Sunda Island and SW Sulawesi ( Govaerts et al. 2015). Some materials from Java have been incorrectly identified as E. winitiana .