Karumballichirus khadroensis

Hyžný, Matúš, Kneer, Dominik & Charbonnier, Sylvain, 2025, The largest ghost shrimps ever: evidence from the fossil record and implications for the maximum size estimate of callianassoid burrowing ghost shrimps, Acta Palaeontologica Polonica 70 (1), pp. 97-113 : 100

Karumballichirus khadroensis (Hyžný & Charbonnier in Hyžný et al., 2016a) comb. nov.

Fig. 2 .

2016a Neocallichirus khadroensis Hyžný & Charbonnier ; Hyžný et al. 2016a: 344, figs. 2, 5A 3, C 2, D 3, 6I.

Material.— Holotype: CPAG.RAN.I.55 (cast MNHN.F.

A52405), left propodus with articulated dactylus; paratypes:

8specimens, CPAG .RAN.I.56–I.63(casts MNHN.F.A52406–

A52413 ), isolated cheliped elements; from Gawar Band section (Khadro Formation, Danian), Ranikot, Sindh, Pakistan .

Description.—Detailed description of the species was provided by Hyžný et al. (2016a) and is therefore not repeated here.

Remarks.— Hyžný et al. (2016a) assumed that the growth rate of Neocallichirus khadroensis was largely the same as the one of Neocallichirus karumba ; the assumption was based on striking morphological similarities between both taxa as shown by Hyžný et al. (2016a: fig. 5). Based on an extrapolation of data on the growth of N. karumba ( Dworschak 2008) , Hyžný et al. (2016a) estimated the total length of the largest individuals of N. khadroensis to be 120 mm. The establishment of Karumballichirus (with the type species N. karumba ) allows re-assignment of N. khadroensis to the respective genus.

Stratigraphic and geographic range.—Danian (Paleocene) of Pakistan ( Hyžný et al. 2016a).

Figures

Fig. 2. Ghost shrimp Karumballichirus khadroensis (Hyžný & Charbonnier in Hyžný et al., 2016a) comb. nov., Danian (Paleocene) of Ranikot, Sindh, Pakistan. A. CPAG.RAN.I.55 (holotype), left major chela (propodus and dactylus) in outer lateral view. B. CPAG.RAN.I.63 (paratype), right major cheliped merus in outer lateral view. C. CPAG.RAN.I.60 (paratype), left major chela (propodus and dactylus) in outer lateral view. All specimens are shown to the same scale.