Aphanocephalus Wollaston, 1873

Yoshitomi, Hiroyuki, 2025, An annotated checklist of the family Discolomatidae (Coleoptera: Coccinelloidea) of the world, Zootaxa 5701 (2), pp. 101-121 : 110-114

Genus Aphanocephalus Wollaston, 1873

Aphanocephalus Wollaston, 1873: 278 .

Type species: Aphanocephalus hemisphaericus Wollaston, 1873 (by monotypy).

Eleothreptus Blackburn, 1895: 235 .

Type species: Eleothreptus punctulatus Blackburn, 1895 (by monotypy). Synonymized by Arrow (1920).

acuminatus Grouvelle, 1918: 55. Seychelles.

aequabilis John, 1956a: 18. Malaysia (Borneo: Sarawak).

africanus Grouvelle, 1912: 212. Tanzania (Kwai, Korogwe).

angulus Pal, 2005: 298. India.

annamensis John, 1956a: 30. Laos, Vietnam, China.

antricolus John, 1964b: 154. Democratic Republic of Congo (Thysville, Grotte, Yangambi).

apoanus John, 1959d: 7. Philippines (Mindanao).

arebiensis John, 1964b: 150. Democratic Republic of Congo (Arebi, Bondo-Mabe).

asoanus John, 1965c: 53. Japan (Kyushu).

assamensis Schawaller, 1989: 50. India.

atrocinctus John, 1942d: 13. Indonesia ( Java).

atomus Grouvelle, 1912: 10. Indonesia ( Sumatra). [ John (1958a) treated it as a junior synonym of A. vitreus , but John (1967a) treated it as a valid species again.]

awashimanus John, 1968b: 15. Japan (Awashima).

banguius John, 1971: 143. Philippines (Luzon).

basilanensis John, 1956a: 12. Philippines ( Basilan).

besucheti Schawaller, 1989: 51. Pakistan.

bimaculatus Grouvelle, 1912: 214. Australia.

= austerus John, 1941a: 194. Australia (Rockhampton). Synonymized by John (1954c).

= bimaculatus Lea, 1921a: 240. Australia ( Queensland). Synonymized by Szawaryn (2025).

= perlucidus John, 1941a: 192. Australia. Synonymized by Szawaryn (2025).

binotatus Grouvelle, 1918: 50. Seychelles.

birmanus Dodero, 1900: 565. Myanmar, China ( Yunnan), Thailand ( Schawaller 1990).

biroi John, 1953a: 79 . Papua New Guinea.

bisbimaculatus John, 1952c: 68. Papua New Guinea.

bisbipunctatus John, 1952c: 67. Papua New Guinea.

blackburni Szawaryn, 2025: 415. Australia ( Queensland). [new name for A. quadrimaculatus Lea, 1921b: 369 ]

blaisei John, 1956a: 14. Vietnam (I. Fai Tsi Long).

bottcheri John, 1971: 143. Philippines ( Basilan).

brevipronotatus John, 1959d: 8. Papua New Guinea (Rabaul).

burgeoni John, 1952e: 49. Democratic Republic of Congo (Haut Uelle, Watsa).

calderi Szawaryn, 2025: 416. Australia ( South Australia).

camerunensis John, 1942d: 16. Cameroon.

castaneus John, 1967a: 118. Papua New Guinea ( Bismarck Archipelago).

chujoi John, 1955b: 53. Japan (Honshu, Shikoku, Kyushu; John 1963b).

claggi John, 1959d: 7. Philippines (Mindanao).

congoensis John, 1956a: 14. Democratic Republic of Congo.

connexus John, 1965e: 292. Democratic Republic of Congo (Yangambi).

convexus Pal, 1996: 12. India.

crinitus John, 1968b: 17. Japan (Ryukyu Isls.).

crucifer John, 1942d: 15. Indonesia ( Java, Sumatra).

cupreus John, 1968c: 192. Indonesia ( Sumatra).

dahli John, 1971: 140. Papua New Guinea ( New Britain).

dasys John, 1955d: 15. Philippines (Luzon).

daulonus John, 1967a: 132. Papua New Guinea.

decellei John, 1964b: 146. Democratic Republic of Congo (Yangambi).

decoomani John, 1954c: 302 . Laos ( Vientiane, Luang Prabang), Vietnam ( Tonkin, Hoa Binh; Schawaller 1993).

dedegwanus John, 1958d: 9. Democratic Republic of Congo.

diffusus John, 1967a: 124. Papua New Guinea.

dilutus John, 1952c: 68. Papua New Guinea.

discrepans John, 1964b: 162. Democratic Republic of Congo (Yangambi).

disjunctus John, 1967a: 130. Papua New Guinea ( New Britain).

distinctus Grouvelle, 1912: 204. Indonesia ( Sumatra).

donisi John, 1964b: 158. Democratic Republic of Congo (Yangambi).

doreyanus John, 1956a: 6. Papua New Guinea.

ebeninus John, 1956a: 8. Vietnam.

fallax John, 1955d: 17 . Taiwan.

feae Dodero, 1900: 566. Myanmar.

formosanus John, 1955d: 17 . Taiwan, Vietnam ( Schawaller 1993).

fruhstorferi John, 1955d: 14. Indonesia ( Sulawesi).

fulgens John, 1967g: 186. Democratic Republic of Congo.

ganganicus Schawaller, 1989: 53. India.

glabrior John, 1955d: 16. Philippines (Luzon).

gorkhus Schawaller, 1987: 4. Nepal.

gracilis John, 1958a: 122. Indonesia ( Sumatra).

gressitti John, 1967a: 128. Papua New Guinea.

guineensis John, 1958a: 120. Papua New Guinea.

habanensis John, 1971: 139. [Locality not given].

hackeri John, 1941a: 191. Australia ( Queensland).

hemisphaericus Wollaston, 1873: 278 . Japan (Honshu, Shikoku, Kyushu).

hilleri John, 1971: 142. Japan.

himalayanus Schawaller, 1989: 53. India, Nepal.

hiranoi Yoshitomi & Pham, 2021a: 90. Vietnam.

hulstaerti John, 1964b: 152. Democratic Republic of Congo ( Equateur, Boteka “Flandria”).

humiphilus John, 1961a: 295. Democratic Republic of Congo.

improvisus John, 1965e: 293. Democratic Republic of Congo (Yangambi).

indicus John, 1956a: 28. India.

infirmus John, 1956f: 71. Democratic Republic of Congo.

infuscatus John, 1956c: 69. Papua New Guinea.

insularis Grouvelle, 1918: 49. Seychelles, Madagascar.

intonsus John, 1956a: 26. Indonesia ( Sumatra).

jesseltonus John, 1967a: 122. Malaysia (Borneo).

johni Pal, 1992: 48 . India.

kalimantanus John, 1959d: 14. Malaysia (Borneo).

kannegieteri John, 1967b: 14. Indonesia ( Java, Sumatra).

kapanganus John, 1964b: 142. Democratic Republic of Congo (Lulua, Kapanga).

kaszabi John, 1962a: 280. Cameroon.

klapperichi John, 1956a: 8. China ( Fujian Province).

kukuyalanji Szawaryn, 2025: 419. Australia ( Queensland).

laeanus John, 1969: 123. Papua New Guinea.

lawrencei Szawaryn, 2025: 421. Christmas Island.

leai Szawaryn, 2025: 422. Australia ( Queensland). [new name for A. quadrinotatus Lea, 1921a: 240 ]

leleupi John, 1964b: 151. Democratic Republic of Congo (Kivu, Kitutu).

lenczyi John, 1962a: 279. Cameroon.

levatus John, 1965e: 293. Democratic Republic of Congo (Yangambi).

levis John, 1956f: 71. Democratic Republic of Congo (Yangambi).

leviterpunctatus John, 1956a: 11. Thailand ( Bangkok).

lewisi John, 1956a: 12. Sri Lanka.

loriai John, 1958a: 118. Papua New Guinea (Woodlark).

loloensis John, 1956a: 16. Cameroon.

maai John, 1967a: 116. Malaysia (Borneo: Sarawak).

mabalianus John, 1964b: 156. Uganda (Tshuapa, Bikoro, Mabali).

macacoanus John, 1964b: 150. Democratic Republic of Congo (Macaco, Luebo).

maculipennis Pal, 2000: 288. India.

masisianus John, 1964b: 154. Democratic Republic of Congo (Kivu, Masisi).

maynei John, 1964b: 160. Democratic Republic of Congo (Yangambi).

mendax John, 1964b: 138. Democratic Republic of Congo (Haut-Uele, Moto).

metallescens Grouvelle, 1912: 203. Indonesia ( Sulawesi).

minutus Grouvelle, 1912: 198. Indonesia ( Sumatra).

modicus John, 1965e: 292. Democratic Republic of Congo (Yangambi).

modiglianii Grouvelle, 1912: 215. Indonesia ( Sumatra).

motoanus John, 1964b: 140. Democratic Republic of Congo (Haut-Uele, Moto).

mutatus John, 1965e: 294. Democratic Republic of Congo (Yangambi).

naius Pal, 2007: 205. Republic of Equatorial Guinea ( Fernando Poo).

niger John, 1954c: 304. India.

nigriparvus John, 1965e: 291. Democratic Republic of Congo (Yangambi).

nigritanus John, 1958a: 117. Papua New Guinea.

nigriventris John, 1955a: 1 . Japan (Honshu, Shikoku, Amami-Oshima; John 1963b).

nitidus Lea, 1922: 307. Australia ( Queensland, New South Wales).

nunukanensis John, 1959d: 12. Malaysia (Borneo).

occidentalis Szawaryn, 2025: 424. Australia ( Western Australia).

pahanganus John, 1967a: 120. Malaysia (Malay Peninsula).

pallidipennis Lea, 1921b: 369. Australia ( Queensland).

= poropterus Lea, 1922: 306. Australia. Synonymized by Szawaryn (2025).

pellitus John, 1956a: 24. Sri Lanka.

pinguis John, 1956a: 12. Indonesia ( Sumatra).

planus John, 1967c: 262. Malaysia (Borneo: Sarawak).

pocsi John, 1965a: 274. Vietnam.

potamophilus Lea, 1921b: 369. Australia (North Western Australia).

= potamophilus John, 1954c: 304. Synonymized by Szawaryn (2025).

praeacutus John, 1956a: 6. Philippines (Mindanao).

prophysus John, 1954c: 303. Singapore.

pseudatomus John, 1954c: 301 . Vietnam ( Tonkin, Hoa Binh).

pubescens Grouvelle, 1912: 207 . Taiwan, Vietnam, Hong Kong, Indonesia ( Sumatra), Malaysia (Borneo, Peninsular; Schawaller 1990, 1993).

ssp. pubescens beccarii John, 1958a: 124. Malaysia (Borneo: Sarawak), Indonesia ( Sumatra), Vietnam.

ssp. pubescens piceus John, 1956a: 9. Indonesia ( Sumatra, Java).

ssp. pubescens sauteri John, 1956a: 9 . Taiwan.

ssp. pubescens schulteissi John, 1956a: 9. Indonesia ( Sumatra, Java).

pulinus John, 1967a: 134. Taiwan.

punctatus Grouvelle, 1912: 200. Indonesia ( Sumatra).

punctipennis Pal, 2008: 243. India (Andaman Is.).

punctulatus ( Blackburn, 1895: 236) . Australia. [orig. comb.: Eleothreptus ]

quadrimaculatus Matthews, 1887: 114 . Malaysia ( Penang), China.

quadrinotatus Grouvelle, 1912: 206 . Indonesia (Kei Isls.).

queenslandicus Szawaryn, 2025: 432. Australia ( Queensland).

quadriplagiatus Grouvelle, 1918: 52. Seychelles.

ranauanus John, 1967a: 126. Malaysia (Borneo: Sabah).

riparius John, 1956c: 401 . Indonesia ( Sumatra).

rufinus John, 1956a: 16. Sri Lanka.

sandakanus John, 1967a: 114. Malaysia (Borneo: Sabah).

satoi John, 1968b: 14. Japan (Tokara Isls.).

saundersi John, 1956a: 22. Singapore, Sri Lanka.

schoutedeni John, 1964b: 141. Democratic Republic of Congo (Mongende).

secretus John, 1956h: 252. Taiwan.

sedlaceki John, 1967c: 260. Solomon Is.

senex John, 1967a: 124. Papua New Guinea.

sexmaculatus John, 1967a: 128. Malaysia (Borneo: Sabah).

shibatai Chûjô & John, 1961: 1. Japan (Amami-Oshima, Tokunoshima, Okinawa-jima).

shirozui John, 1958b: 61 . Japan (Amami-Oshima).

shompen Pal, 2008: 244. India (Nicobar Is.).

sikkima Pal, 1996: 10. India.

silvanus John, 1965e: 291. Democratic Republic of Congo (Yangambi).

similis John, 1964b: 160. Democratic Republic of Congo (Yangambi).

slipinskii Szawaryn, 2025: 433. Australia ( New South Wales, Queensland).

splendens Grouvelle, 1912: 201. Indonesia ( Sumatra).

ssp. splendens hirashimai John, 1967a: 118. Malaysia (Borneo: Sabah).

subdepressus Grouvelle, 1918: 53. Seychelles.

submendax John, 1964b: 148. Democratic Republic of Congo (Haut-Uele, Moto).

superbus John, 1967a: 122 . Malaysia (Borneo: Sabah).

tabaci John, 1956a: 28. Indonesia ( Sumatra), Papua New Guinea.

takaoensis nom. nov. Taiwan. [new name for A. sauteri John, 1971: 139 ].

taiwanus John, 1967a: 134. Taiwan.

takeii John, 1963a: 8. Japan (Honshu, Shikoku, Kyushu, Tsushima). [as subspecies of A. nigriventris ]

tamaii John, 1968b: 18. Japan (Ryukyu Isls.).

tetrasignatus John, 1952c: 69. Papua New Guinea.

tonkinensis John, 1954c: 302 . China ( Yunnan), Vietnam ( Tonkin, Hoa Binh).

tsushimanus John, 1961c: 1. Japan (Tsushima).

turbidus John, 1967g: 187. Democratic Republic of Congo.

uelensis John, 1964b: 142. Democratic Republic of Congo (Haut-Uele, Watsa).

ugandanus John, 1960b: 319. Uganda, Democratic Republic of Congo, Angola ( John 1966b).

uncus John, 1964b: 163. Democratic Republic of Congo (Yangambi).

ventricosus John, 1958a: 116. Indonesia ( Sumatra).

vitreus Matthews, 1887: 5 . China, Malaysia (Borneo: Sabah, Sarawak; Peninsular; Schawaller 1990, 1993), Indonesia ( Java, Sumatra), Papua New Guinea.

weiri Szawaryn, 2025: 436. Australia ( Queensland).

wollastoni Rye, 1873: 167. Japan (Honshu, Kyushu, Okinawa-jima, Ishigaki-jima, Iriomote-jima), Vietnam ( Schawaller 1993).

yangambiensis John, 1959d: 6. Democratic Republic of Congo (Yangambi).