Tomosvaryella caesariata Motamedinia, Skevington & Földvari, 2023

Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, Zootaxa 5599 (1), pp. 1-271 : 54

publication ID	https://doi.org/10.11646/zootaxa.5599.1.1
publication LSID	lsid:zoobank.org:pub:B140A7ED-4B89-464B-8A3E-16934B175A40
DOI	https://doi.org/10.5281/zenodo.14971076
persistent identifier	https://treatment.plazi.org/id/715487A7-FFE7-EC5B-D8D9-EFF6FC4E17E0
treatment provided by	Plazi
scientific name	Tomosvaryella caesariata Motamedinia, Skevington & Földvari
status	sp. nov.

Treatment

Tomosvaryella caesariata Motamedinia, Skevington & Földvari sp. nov.

urn:lsid:zoobank.org:act:C6E322C9-C034-4E07-85A2-F5AA04CC5825

Figs 21A–E View FIGURE 21 , 112 View FIGURE 112 , 145C View FIGURE 145 , 152B View FIGURE 152

Diagnosis: This species can be recognized by dense long bristles covering the abdominal tergites ( Fig. 152B View FIGURE 152 ); hind trochanter with two patches of bristles ventroapically and ventrobasally ( Fig. 145C View FIGURE 145 ); elongated surstyli in dorsal view ( Fig. 21A View FIGURE 21 ); gonopods with two thumb-like projections extended towards surstyli, one of the ejaculatory ducts with some teeth in basal half in ventral view ( Fig. 21B View FIGURE 21 ).

Description: MALE: Body length: 4.0– 4.2 mm. Head. Scape and pedicel brown, Flagellum tapering, yellow-brown. Face silvery-yellow pollinose. Frons, upper part shining black; lower part distinctly silver-yellow pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, covered by scattered short light bristles. Thorax. Postpronotum pale, light brown pollinose, with 4–5 long dark bristles along upper margin. Mesonotum (viewed obliquely from front) light brown pollinose, with long bristles at anterior supra-alar area in a row, extended to posterior and uniseriate rows of long intra-alar bristles. Scutellum silvery brown pollinose and with 24–26 long bristles along distal edge. Halter, knob, stem pale, tip brownish, stem with three short bristles dorsally. Legs. Fore coxa gray pollinose with 10 short brown bristles in a row. Mid coxa with 3–4 long dark bristles and 2–3 short light brown bristles. Coxae, trochanters, femora, tibiae dark brown, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/6 of tibiae yellow (tibiae otherwise dark brown), tarsal segments brown. Hind trochanter with two patches of condensed spine-like dark bristle ventrobasally and ventroapically ( Fig. 145C View FIGURE 145 ); ventrobasal spines present on fore and mid femora. Fore and mid femora with some short bristles posteroventrally in basal half. Hind femur without ventroapical spines, except 8–12 equally spaced bristles posteroventrally in distal half. Subapical (distal) spines on first four tibiae present (short). Hind metatarsus distinctly flattened, extended toward one side, covered by erect bristles especially along lateral edge. Pulvilli shorter than last tarsal segment. Wing. Length: 3.9–4.1 mm. light brown color. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1–3 long dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on tergite 5 larger). All tergites covered by condensed long strong brown bristles as long as width of hind femur at distal end ( Fig. 152B View FIGURE 152 ). Lateral bristles on first tergite present, 32–38 long dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: T6, S7 invisible; T5 0.4–0.5 times as long as ST8. Genitalia without dissection: ST8 grey pollinose, covered by long bristles, round, globular in dorsal view, brown; membranous area is slit-like, mostly directed posteriorly, extended close to epandrium; Genitalia. Genital capsule in dorsal view: epandrium brown without pollinosity, surstyli brown; epandrium wider at base. Surstyli widened at base, elongated, slightly curved to each other at apex, left surstylus is longer ( Fig. 21A View FIGURE 21 ). Genital capsule in ventral view: both gonopods equal in height with a thumb-like projection extended towards surstyli, rounded at apex, with a distinct projection in their inner margins before apex; subepandrial sclerite distinct, with a fin-shaped projection in its center, phallus trifid, one of ejaculatory ducts with some teeth at the basal third; phallic guide with 3–5 spines dorso-dorsolaterally, one is longer ( Fig. 21B View FIGURE 21 ); Genital capsule in lateral view: left surstylus straight in basal two thirds, bent toward sternite in apical thirds, right surstylus short and rather straight ( Fig. 21D–E View FIGURE 21 ). Ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 21C View FIGURE 21 ).

FEMALE: Unknown.

Etymology: From the Latin caesariatus, covered with hair or long-haired, in reference to the densely haired abdomen of this species.

Examined material: HOLOTYPE: AUSTRALIA: Queensland: 3.5 km SW by S Mount Baird , 15°10’S, 145°7’E, 3.V.1981, D.H. Colless, Malaise trap, JSS8580 (1♂, ANIC) GoogleMaps ; PARATYPES: AUSTRALIA: Queensland: 3.5 km SW by S Mount Baird , 15°10’S, 145°7’E, 4. V GoogleMaps .1981, D.H. Colless, Malaise trap, JSS199 (1♂, ANIC); Brisbane, [27°28’S, 153°1’E] GoogleMaps , 23.VII .1964 , I.C. Yeo, JSS8266 (1♂, QM).

Distribution: Australia (Queensland) ( Fig. 112 View FIGURE 112 ).

Notes: The distribution of this species is suspicious, and the old Brisbane specimen could be mislabeled. It is strange that no other specimens have been found around Brisbane with the intensity of collecting there. More work should be conducted in the type locality to determine habitat preferences.