Tomosvaryella brooksi Motamedinia, Skevington & Földvari, 2023

Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, Zootaxa 5599 (1), pp. 1-271 : 50

Tomosvaryella brooksi Motamedinia, Skevington & Földvari sp. nov.

urn:lsid:zoobank.org:act:DFC7F93C-2B7A-46E8-BE5E-297E5C7609E2

Figs 19A–E , 115

Diagnosis: This species can be recognized by the shape of surstyli, both are triangle-shaped and the right surstylus is wider than left one in the middle in dorsal view ( Fig. 19A ); gonopods asymmetrical, right is longer than left one, both with a short semicircular projection at the apex; each of ejaculatory ducts with 2–3 small spines in ventral view ( Fig. 19B ); both surstyli straight and widened, pointed at apex in lateral view ( Fig. 19D–E ).

Description: MALE: Body length: 2.5 mm. Head. Scape, pedicel and arista dark brown, Flagellum tapering, yellow-brown, light brown pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.2 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale, with 3–4 long pale bristles. Prescutum with two triangle-shaped grey marks on anterior part. Mesonotum (viewed obliquely from front) silvery-brown pollinose. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Scutellum silvery brown pollinose and with 6–8 scattered bristles. Halter, base dark, stem pale, knob pale, tip brownish. Legs. Mid coxa with 3–4 dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright yellow completely, except last tarsomeres. Hind trochanter triangle-shaped grey pollinose without specific feature; ventrobasal spines absent on fore and mid femora. Hind femur without ventroapical spines, except 6–10 equally spaced short bristles posteroventrally and anteroventrally, longest up to 1/2 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2–4 combined, dorsally all tarsomeres with erect pale bristles except last tarsomeres with longer dark bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.4 mm. Upper side of basal costal cell with one long light brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2–3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, all tergite silvery-brown pollinose, Dispersed short brown bristles on all tergites present. Lateral bristles on first tergite present, 3–6 dark bristles up to as long as half of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: membranous area round and wide, epandrium longer than wide (MLE: MWE = 1.3). Surstyli triangle-shaped, wide at the base, pointed to each other at apex, right surstylus is wider than left one in the middle, left one is longer ( Fig. 19A ). Genital capsule in ventral view: gonopods asymmetrical, right is longer than left one, both with a short semicircular projection at the apex, small hypandrium, long subepandrial sclerite; phallic guide long; phallus trifid, each one with 2–3 small spines ( Fig. 19B ); Genital capsule in lateral view: both surstyli straight and widened, pointed at apex ( Fig. 19D–E ). Ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 19C ).

FEMALE: Unknown.

Etymology: This species is named in honour of Scott E. Brooks. Scott collected one of the two known specimens and is a well-known dipterist and colleague of the authors. Scott works at the CNC in Ottawa, Ontario, Canada.

Examined material: HOLOTYPE: AUSTRALIA: Queensland: Carnarvon Nat [ional] Park, Consuelo land, Race Course Spring , 24°56’S, 148°5’E, Vegetation, 12.X.2002, S. Boucher, sweep net, LEM _0016155 (1♂, QM) ; PARATYPE: AUSTRALIA: Queensland: Carnarvon National Park, The Tombs , 25°5’S, 147°52’E, 12.X.2002, S. E. Brooks, sweep net, LEM _ 0016200 (1♂, LEM) .

Distribution: Australia (Queensland) ( Fig. 115 ).

Notes: This species is only known from two specimens collected in Carnarvon National Park in Queensland. The species presumably does not hilltop as there has been a lot of effort collecting on hilltops in Carnarvon and it has not appeared in any of those samples.

Figures

FIGURE 19. Male genitalia of Tomosvaryella brooksi sp. nov. (LEM_0016155, QM) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view.

FIGURE 115. Distribution of Tomosvaryella brooksi sp. nov., T. petala sp. nov., T. planca sp. nov. in Australia.