Tomosvaryella arnhemensis Motamedinia, Skevington & Földvari, 2023

Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, Zootaxa 5599 (1), pp. 1-271 : 26

publication ID	https://doi.org/10.11646/zootaxa.5599.1.1
publication LSID	lsid:zoobank.org:pub:B140A7ED-4B89-464B-8A3E-16934B175A40
DOI	https://doi.org/10.5281/zenodo.14971016
persistent identifier	https://treatment.plazi.org/id/715487A7-FFCB-EC77-D8D9-EFF6FF5D1624
treatment provided by	Plazi
scientific name	Tomosvaryella arnhemensis Motamedinia, Skevington & Földvari
status	sp. nov.

Treatment

Tomosvaryella arnhemensis Motamedinia, Skevington & Földvari sp. nov.

urn:lsid:zoobank.org:act:8EEC7262-DAF5-4439-B49B-683CD1989B4F

Figs 8A–E View FIGURE 8 , 109 View FIGURE 109 , 145A View FIGURE 145

Diagnosis: This species can be recognized by hind trochanter with 6–10 short dark bristles in a row from ventrobasally to ventroapically ( Fig. 145A View FIGURE 145 ); tiny surstyli, upright in dorsal view ( Fig. 8A View FIGURE 8 ); gonopods extended towards surstyli, rounded at apex with a distinct projection on inner margin; phallic guide with 2–4 short dorso- dorsolateral spines, one of ejaculatory ducts with some short teeth in basal half in ventral view ( Fig. 8B, D–E View FIGURE 8 ).

Description: MALE: Body length: 2.7 mm. Head. Scape, pedicel dark brown, flagellum acuminate, yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1/3. Thorax. Postpronotum yellow, with 3–4 pale bristles. Mesonotum (viewed obliquely from front) silvery brown pollinose, anterior part without pollinosity. Dorsocentral bristles short, pale, numerous pale bristles around postpronotum present. Scutellum silvery brown pollinose and with 4–8 short dark bristles. Halter, knob pale, tip brownish, stem brown. Legs. Mid coxa with 3–4 long dark bristles. Fore and mid trochanter with two short dark bristles ventroapically; trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright brown. Hind trochanter with 6–10 short dark bristles in a row from ventrobasally to ventroapically ( Fig. 145A View FIGURE 145 ); ventrobasal spines present (2) on fore and mid femur. Hind femur without ventroapical spines, except 8–10 equally spaced bristles posteroventrally. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres gently flattened (t1–3), hind metatarsus almost as long as 2–4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 silvery grey. Dispersed short dark brown bristles on all tergites present. Lateral bristles on first tergite present, 3–5 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE:MWE = 0.7). Surstyli tiny (rather hard to see in dorsal view), upright, widened at the base, angled straight upwards in apical half ( Fig. 8A View FIGURE 8 ). Genital capsule in ventral view: both gonopods equal in height, extended towards surstyli, rounded at apex, gonopods with a distinct projection on inner margin; phallic guide with 2–4 short dorso- dorsolateral spines; phallus trifid, one of ejaculatory ducts with some teeth in basal half; subepandrial sclerite distinct ( Fig. 8B View FIGURE 8 ); Genital capsule in lateral view: both surstyli straight in basal two thirds, bent in apical third ( Fig. 8D–E View FIGURE 8 ). Ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 8C View FIGURE 8 ).

FEMALE: Unknown.

Etymology: This species name refers to Arnhem Land, a historical region of the Northern Territory of Australia where the only known specimen was collected.

Examined material: HOLOTYPE: AUSTRALIA: Northern Territory: Arnhem Land, Maningrida , 12°3’S, 134°13’E, 5m, 20–21.III.1961, L.&M. Gressitt, CNCD5563 (1♂, BPBM). GoogleMaps

Distribution: Australia (Northern Territory) ( Fig. 109 View FIGURE 109 ).

Notes: This species is known from a single specimen from a coastal area near a river mouth in the Northern Territory. Based on this limited information, it should be looked for in either mangroves or coastal monsoon forest.