Tomosvaryella hardyi Motamedinia, Skevington & Földvari, 2023

Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, Zootaxa 5599 (1), pp. 1-271 : 103-104

Tomosvaryella hardyi Motamedinia, Skevington & Földvari sp. nov.

urn:lsid:zoobank.org:act:4DE49584-1E6A-47A8-B4A4-73DF92F3EAED

Figs 45A–D , 122

Diagnosis: This species can be recognized by elongated surstyli with two distinct triangle-shaped projections at the inner margin before the apex in dorsal view ( Fig. 45A ); gonopods equal in height, extended towards surstyli, right gonopod wider with three dentations at apex in ventral view ( Fig. 45B ); phallus with a small triangular projection before emerging ejaculatory ducts in lateral view ( Fig. 45C–D ).

Description: MALE: Body length: 1.1 mm (without head and abdomen). Thorax. Postpronotum light yellow, with 2–3 short pale bristles. Mesonotum (viewed obliquely from front) silvery-brown pollinose; scutum with scattered dark bristles at anterior supra-alar area, extended to posterior. Postalar callus dark brown with 1–2 short light brown bristles in the middle. Scutellum silvery pollinose and with a few (3–5) short dark bristles along the distal edge. Halter, knob pale, stem light brown, base dark. Legs. Mid coxa with 2–3 dark bristles. Coxae, trochanters, femora dark, hind femur shining ventrally, knees and basal 1/3 of tibiae light brown (tibiae otherwise dark brown), tarsomeres brown. Fore and mid trochanter with two short dark bristles; hind trochanter silvery pollinose covered by scattered short light brown bristles; two ventrobasal spines absent on fore mid femur; two ventroapical spines present on fore mid femur. Hind femur with two rows short spines ventrally. Hind tibia with two rows of short brown bristle on anterior and posterior side. Hind tarsomeres not flattened, covered by short light brown bristles ventrally and short brown bristles dorsally, hind metatarsus as long as 2–4 combined; pulvilli shorter than last tarsal segment. Wing. (one wing missing). Length: 3.1 mm. Upper side of basal costal cell with two short dark brown bristles and two same-sized light brown bristles. Fourth costal section 1.5–2 times as long as third costal section. Cross-vein r-m at middle of discal cell. Abdomen. Tergites 1 silvery pollinose with 5–6 different-sized lateral bristles; Genitalia. Genital capsule in dorsal view: epandrium slightly longer than wide (MLE:MWE = 1.1). Surstyli symmetrical, elongated, widened at base, moderately narrowed before apex, with two distinct triangle-shaped projections at inner margin before apex ( Fig. 45A ). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, right gonopod wider with three dentations at apex ( Fig. 45B ); phallic guide strong; subepandrial sclerite small and wide ( Fig. 45B ); Genital capsule in lateral view: both surstyli widened at the base, narrowed and straight at the middle, then bent towards sternite, phallus with a small triangular projection before emerging ejaculatory ducts ( Fig. 45C–D ).

FEMALE: unknown.

Examined material: HOLOTYPE: AUSTRALIA: Queensland: Cairns , 16°55’S, 145°46’E, VIII.1904, CNCD5550 (1♂, BPBM)

Etymology: This new species is named in honor of D. Elmo Hardy, who discovered it as a new species but never published a description.

Distribution: AUSTRALIA: (Queensland) ( Fig. 122 ).

Notes: The sole specimen of this species is a syntype of T. synadelpha but Hardy recognized it was different from the others and likely a new species.

Figures

FIGURE 45. Male genitalia of Tomosvaryella hirta sp. nov. (JSS8906, QM) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view.

FIGURE 122. Distribution of Tomosvaryella dissidens sp. nov., T. gidgee sp. nov., T. hardyi sp. nov. in Australia.