Tomosvaryella gaimarii Motamedinia, Skevington & Földvari, 2023

Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, Zootaxa 5599 (1), pp. 1-271 : 97-99

publication ID	https://doi.org/10.11646/zootaxa.5599.1.1
publication LSID	lsid:zoobank.org:pub:B140A7ED-4B89-464B-8A3E-16934B175A40
DOI	https://doi.org/10.5281/zenodo.14971164
persistent identifier	https://treatment.plazi.org/id/715487A7-FFB2-EC0C-D8D9-EB03FBD712B8
treatment provided by	Plazi
scientific name	Tomosvaryella gaimarii Motamedinia, Skevington & Földvari
status	sp. nov.

Treatment

Tomosvaryella gaimarii Motamedinia, Skevington & Földvari sp. nov.

urn:lsid:zoobank.org:act:953F0804-AFEB-43E4-8D04-DFECA8C7A7DE

Figs 42A–E View FIGURE 42 , 123 View FIGURE 123 , 144B View FIGURE 144

Diagnosis: This species can be recognized by the two distinct spines on the hind trochanter ( Fig. 144B View FIGURE 144 ); abdominal sternites 3–4 with some spine-like bristles along posterior margin; surstyli slender and long in dorsal view ( Fig. 42A View FIGURE 42 ); gonopods widened with a distinct lobe towards each other in ventral view ( Fig. 42B View FIGURE 42 ); two ejaculatory ducts with spines ( Fig. 42B, D View FIGURE 42 ); surstyli with a distinct triangle-shaped lobe dorsobasally in lateral view ( Fig. 42D–E View FIGURE 42 ).

Description: MALE: Body length: 2.6 mm. Head. Scape, pedicel and Flagellum dark brown; scape with two short upper bristles. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery-brown pollinose with scattered pale short bristles. Thorax. Postpronotum pale, with 4–5 long pale bristles. Mesonotum (viewed obliquely from front) silvery-brown pollinose, with some longer supra-alar bristles behind postpronotum. Scutellum brown pollinose and with 4–6 bristles along distal edge. Halter, knob pale, tip yellow, stem brown. Legs. Mid coxa with 3–4 long brown bristles. Trochanters brown, femora dark brown, shining ventrally; knees and basal 1/6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright brown. Fore coxa with three short pale bristles ventromedially. Hind trochanter with two distinct spines ( Fig. 144B View FIGURE 144 ), ventromedial one darker, posteroventral one lighter and longer, hook like shape; ventrobasal spines present (1) on fore and absent on mid femur. Hind femur without ventroapical spines, two rows short light bristles posteroventrally. Fore femur with two distinct and long bristles ventrobasally. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres distinctly flattened except last one, hind metatarsus almost as long as 2–4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one long light brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3–5 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, covered with short light brown bristles, tergite 1 brown-silvery grey. Lateral bristles on first tergite present, 4–5 dark bristles up to as long as ¾ of hind femur’s width at base. Sternite 3–4 with 4–6 short spine-like bristles along posterior margin. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE:MWE = 0.5). Surstyli slender and elongated, curved to each other at apex, left surstylus is little longer ( Fig. 42A View FIGURE 42 ), both surstyli with a distinct lobe at the base. Genital capsule in ventral view: both gonopods extended towards surstyli and widened, pointed at apex with a distinct lobe on inner margin towards each other; phallic guide long, pointed at apex; phallus with three ejaculatory ducts, two ejaculatory ducts with spines, one spine is longer; subepandrial sclerite long ( Fig. 42B View FIGURE 42 ); Genital capsule in lateral view: both surstyli gently curved towards sternite, both with a distinct triangle-shaped lobe dorsobasally ( Fig. 42D–E View FIGURE 42 ). Ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 42C View FIGURE 42 ).

FEMALE: Unknown.

Etymology: Named in honour of Stephen D. Gaimari, one of the collectors of the only known specimen and a major contributor to Diptera research.

Examined material: HOLOTYPE: AUSTRALIA: Queensland: Brisbane Forest Park, top of Scrub Road , 27°25’S, 152°51’E, 370m, sclerophyll Eucalyptus forest, 1–7.XII.1995, Irwin , Gaimari , JSS16633 (1♂, QM). GoogleMaps

Distribution: Australia (Queensland) ( Fig. 123 View FIGURE 123 ).

Notes: It is surprising that only a single specimen of this species has been found as it is in a well-collected area. Different people have run long term Malaise trapping programs here and there has been a lot of hand collecting as well. The area contains a mixture of dry sclerophyll and wet sclerophyll (vine) forest.