Tomosvaryella parkeri Földvari, Skevington & Motamedinia, 2023

Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, Zootaxa 5599 (1), pp. 1-271 : 161

publication ID	https://doi.org/10.11646/zootaxa.5599.1.1
publication LSID	lsid:zoobank.org:pub:B140A7ED-4B89-464B-8A3E-16934B175A40
DOI	https://doi.org/10.5281/zenodo.14971294
persistent identifier	https://treatment.plazi.org/id/715487A7-FF72-ECCE-D8D9-EE2BFD5417A8
treatment provided by	Plazi
scientific name	Tomosvaryella parkeri Földvari, Skevington & Motamedinia
status	sp. nov.

Treatment

Tomosvaryella parkeri Földvari, Skevington & Motamedinia sp. nov.

urn:lsid:zoobank.org:act:5BFE6F07-15FE-4F5F-B6DF-7FB80DBE9136

Figs 72A–E View FIGURE 72 , 111 View FIGURE 111 , 155C View FIGURE 155

Diagnosis: This species can be recognized by elongated surstyli that are moderately narrowed to the apex ( Fig. 72A View FIGURE 72 ); unequal gonopods, right gonopod shorter with two triangle-shaped projections on inner margin in ventral view ( Fig. 72B View FIGURE 72 ); distinct and long hypandrial apodeme ( Fig. 72B View FIGURE 72 ); phallic guide with 3–5 dorsolateral spines ( Fig. 72B, D–E View FIGURE 72 ).

Description: MALE: Body length: 3.1 mm. Head. Flagellum short acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1/3. Thorax. Postpronotum pale, with 2–3 pale bristles. Mesonotum (viewed obliquely from front) grey pollinose, anterior part more intensely pollinose, grayish also from the side. Scutellum silvery pollinose and with 6–8 bristles along the distal edge. Dorsocentral bristles short, dark, somewhat longer along frontal edge, numerous pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with two dark bristles, one longer than the other. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/6 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow-brown dorsally, ventrally yellow, last segment darker. Hind trochanter with a half circle (flat part dorsally), no distinct bristles; ventrobasal spines absent on fore and mid femur. Ventroapical row of spines on fore femur absent; 4–5 small, black, spines on mid femur; hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae absent. Hind metatarsus distinctly flattened (tarsomeres 1–3 most flattened), first tarsomere slightly longer than 2–4 combined, with scrub-like bristles ventrally (tarsomeres 1–2 only). Pulvilli shorter than last tarsal segment. Wing. Length: 3.2 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2.2 times as long as third costal section. Cross-vein r-m distinctly distal to middle of discal cell. 5–6 distinct brown setulae on tegula. Abdomen. Dissected. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE:MWE = 1.4). Surstyli rather symmetrical, elongated, moderately narrowed to apex, left surstylus longer than right one, right surstylus bent to left one before apex ( Fig. 72A View FIGURE 72 ). Genital capsule in ventral view: gonopods unequal in height, left is longer, right is wider, right gonopod with two different-sized triangular projections on inner margin, left gonopod with one smaller triangular projection; distinct, long hypandrial apodeme; phallic guide with 3–5 dorso- dorsolateral spines, one is longer, distinct subepandrial sclerite ( Fig. 72B View FIGURE 72 ); Genital capsule in lateral view: both surstyli broadened at base, moderately narrowed to apex, curved towards sternite, phallic guide strong, pointed towards sternite at apex ( Fig. 72D–E View FIGURE 72 ); ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 72C View FIGURE 72 ).

FEMALE: Body length: 2 mm (without head). As male except for the following characters. Frons, eyes: NO HEAD. Pulvilli and claws about 2–2.8 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergite 5 only. Ovipositor. Straight in ventral view, strongly bent (90 degrees at the base of piercer) in lateral view (yellow piercer, brown base), reaching proximal end of 5th segment; base silvery pollinose with occasional bristles. LP:LB = 2.4. LDP:LPP = 4.0. ( Fig. 155C View FIGURE 155 ).

Etymology: This species is named in honour of Frank D. Parker who collected both known specimens with Mike Irwin. Frank was involved in the collection of 6% of the specimens and 15% of the species used in this paper (Supplementary file 2).

Examined material: HOLOTYPE: AUSTRALIA: Western Australia: 47 km S Pardoo R [oa]d House on Shay Gap R [oa]d, 20°23’S, 120°1’E, 170m, dry wash n[ea]r Spinifex , 1–14.V.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15702 (1♂, WAM) GoogleMaps ; PARATYPE: AUSTRALIA: Western Australia: same data as holotype, JSS15704 (1♀, QM) GoogleMaps .

Distribution: Australia (Western Australia) ( Fig. 111 View FIGURE 111 ).

Note: Intraspecific genetic distance is 0.0%. This species is genetically most similar to T. quadrata sp. nov. (7.5% pairwise divergence) (Supplementary file 3).