Tomosvaryella patula Földvari, Skevington & Motamedinia, 2023
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publication ID |
https://doi.org/10.11646/zootaxa.5599.1.1 |
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publication LSID |
lsid:zoobank.org:pub:B140A7ED-4B89-464B-8A3E-16934B175A40 |
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DOI |
https://doi.org/10.5281/zenodo.14971304 |
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persistent identifier |
https://treatment.plazi.org/id/715487A7-FF71-ECCA-D8D9-EA77FD4F1770 |
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treatment provided by |
Plazi |
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scientific name |
Tomosvaryella patula Földvari, Skevington & Motamedinia |
| status |
sp. nov. |
Tomosvaryella patula Földvari, Skevington & Motamedinia sp. nov.
urn:lsid:zoobank.org:act:9F645DC0-660C-4972-92BE-FBA870D01E99
Figs 74A–E View FIGURE 74 , 134 View FIGURE 134 , 149D View FIGURE 149 , 155D View FIGURE 155
Diagnosis: This species can be recognized by the hind trochanter having a keel covered by short dark spines on the edge ( Fig. 149D View FIGURE 149 ); hind metatarsus distinctly flattened; elongated surstyli, left surstylus with a lobe on inner margin in dorsal view ( Fig. 74A View FIGURE 74 ); both gonopods with a triangular projection on inner margin; phallic guide with 3–5 dorsolateral spines ( Fig. 74B, D–E View FIGURE 74 ).
Description: MALE: Body length: 2.9–3.1 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1/3. Thorax. Postpronotum pale, with 3–4 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 6–8 bristles along the distal edge. Dorsocentral bristles short, dark, somewhat longer along frontal edge, numerous pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 2–3 long dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow, ventrally paler, last segment brown. Hind trochanter with shiny edge (keel) at 45 degrees to the long axis of trochanter, with short dark spines on the edge ( Fig. 149D View FIGURE 149 ); weak ventrobasal spines (2) present on fore and missing on mid femur. Ventroapical row of spines on fore femur absent; 4–5 small, black spines on mid femur; hind femur without ventroapical spines, except 8–10 equally spaced bristles on distal half posteroventrally on a widened femur in the middle (almost like a small keel), longest up to 1.5 times the width of hind tibia at distal end (sometimes adpressed to femur). Subapical (distal) spines on first four tibiae present (short). Hind metatarsus distinctly flattened, as long as 2–5 combined and with strong bristles dorsally, scrub-like bristles ventrally (tarsomeres 1–2 only). Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2–2.5 times as long as third costal section ( Fig. 158E View FIGURE 158 ). Cross-vein r-m at middle of discal cell (sometimes slightly distal to middle). 2–3 short brown setulae on tegula. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on t5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1/4 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4–6 short dark bristles up to as long as 2/3 of hind femur’s width at base. Postabdomen in dorsal view: T6, S7 invisible; T5 0.5–0.7 times as long as ST8. Genitalia without dissection: ST8 large, rounded in dorsal view, brown and with a few distinct bristles, otherwise with velvet-like coverage; membranous area with round part extended dorsally and narrow part approaching epandrium, mostly directed posteriorly; epandrium brown, surstyli yellow. Genitalia. Genital capsule in dorsal view: epandrium slightly as long as wide (MLE:MWE = 1.08). Surstyli elongate, left surstylus with a distinct lobe on inner margin in middle and longer than right one, right surstylus with a small denticle on inner margin after basal fourth, both surstyli strongly bent to each other at apex ( Fig. 74A View FIGURE 74 ). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, both gonopods with a triangular projection on inner margin; phallic guide with 3–6 dorso-dorsolateral spines, one is longer, distinct subepandrial sclerite ( Fig. 74B View FIGURE 74 ); Genital capsule in lateral view: both surstyli broadened at base, rather straight, right surstylus moderately narrowed to apex, left surstylus bent towards sternite before apex ( Fig. 74D–E View FIGURE 74 ); ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 74C View FIGURE 74 ).
FEMALE: Body length: 2.4 mm (without head). As male except for the following characters. Frons, eyes: NO HEAD. Pulvilli and claws about 2.5–3 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4–6. Ovipositor. Straight in ventral view, straight and robust in lateral view (yellow piercer, brown base), reaching middle of 3 rd segment; base silvery pollinose with dispersed bristles. LP:LB = 3.0. LDP:LPP = 5.0. ( Fig. 155D View FIGURE 155 ).
Etymology: From the Latin patulus, spread out, broad, in reference to the distinctly flattened hind metatarsus.
Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Warrumbungle National Park , 31°14’S, 149°1’E, 19.X–3.XI.1997, J. Skevington, & S. Winterton, Malaise trap, JSS8439 (1♂, QM) GoogleMaps ; PARATYPES: AUSTRALIA: New South Wales: Colo Heights, Putty Road, 33°23’S, 150°45’E, 6.II.1968, D.H. Colless, JSS8667 (1♂, ANIC); Culgoa National Park , 13.5km SW Cawwell Homestead ( CGN1 View Materials M), 29°7’S, 146°56’E, Callitris , 30.I– 18. V GoogleMaps .2010, QM GoogleMaps Team, B. Shiebaan & R . Ohlsen, Malaise trap, CNC576617 View Materials (1♂, QM); Culgoa National Park, 8km West-Northwest Cawwell Homestead, Diemunga Lagoon ( CGN2 View Materials M), 29°3’S, 146°60’E, Coolibah, 30.I–18. V GoogleMaps .2010, C. Lambkin & R . Olsen, B. Shieban, Malaise trap, 19322, CNC575139 View Materials (1♂, QM); Bundjalung National Park; near Evans Head , 29°2’S, 153°26’E, 8.I.1997, C.L. Lambkin, Malaise trap, JSS6068 (1♀, QM) GoogleMaps ; Queensland: Carnarvon National Park, Mount Moffatt Section, The Tombs , 25°5’S, 147°52’E, 10.X.2002, J. Skevington, hand collected, JSS12635 (1♂, CNC); Carnarvon National Park , Mount Moffatt Summit , 25°4’S, 148°3’E, 1097m, hilltop, 2. XII GoogleMaps .1997, J. Skevington & C. Lambkin, hand collected, JSS1821 (1♂, QM); Carnarvon National Park, Mount Moffatt Summit, 25°4’S, 148°3’E, 1097m, hilltop, 13.X.2002, J. Skevington, JSM 466, CNC484088 View Materials (1♀, CNC); Isla Gorge National Park , 25°11’S, 149°58’E, 320m, hilltop near lookout, 13.XI.1999, J.&A. Skevington, hand collected, JSS7256 (1♂, QM); Western Australia : West of Norseman , 32°11’S, 121°38’E, Eucalyptus woodland , 1–17.XI.2003, C. Lambkin & J. Recsei, Malaise trap, JSS16212 (1♂, ANIC) GoogleMaps .
Distribution: Australia (New South Wales, Queensland, Western Australia) ( Fig. 134 View FIGURE 134 ).
Notes: Intraspecific genetic distance ranges from 0.3% to 1.5%. This species is genetically most similar to T. unda sp. nov. (1.6–8.3% pairwise divergence) (Supplementary file 3). This is a forest species that is apparently widespread but rare. It has been found hilltopping.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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