Tomosvaryella prolata Motamedinia, Skevington & Földvari, 2023

Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, Zootaxa 5599 (1), pp. 1-271 : 177-178

Tomosvaryella prolata Motamedinia, Skevington & Földvari sp. nov.

urn:lsid:zoobank.org:act:F99AFE16-6998-476A-9475-6405D9E5DEDA

Figs 80A–E , 135

Diagnosis: This species can be recognized by the elongated surstyli, moderately narrowed towards the apex in dorsal view ( Fig. 80A ); equal gonopods in height, extended towards surstyli; phallic guide with 2–4 different-sized spines dorsolaterally, subepandrial sclerite long, rounded in middle in ventral view ( Fig. 80B ).

Description: MALE: Body length: 2.0 mm (without abdomen). Head. Scape and pedicel dark brown. Pedicel with 2–4 upper bristles; flagellum brown and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput grey pollinose, covered by short brown bristles. Thorax. Mesonotum (viewed obliquely from front) silvery-brown pollinose; scutum with scattered bristles on anterior supra-alar area. Postalar callus dark brown with 2–3 short bristles in the middle. Scutellum silvery pollinose and with a few indistinct bristles along the distal edge. Halter, knob pale, stem light brown, base dark. Legs. Coxae, trochanters, femora dark, hind femur shining ventrally, knees and basal ¼ of tibiae yellow (tibiae otherwise dark brown), tarsomeres light brown. Hind trochanter silvery pollinose, with some short bristles in a row; ventrobasal spines absent on fore and mid femur. Hind femur with two rows of ventroapical spines. Hind tibia with two rows of short black bristles on anterior and posterior side. Hind tarsomeres moderately flattened, 5th hind tarsomere as long as 3–4 combined; pulvilli shorter than 5th tarsal segment. Wing. Length: 2.5 mm. Fourth costal section 1.5–2 times as long as third costal section. Cross-vein r-m at middle of discal cell. Abdomen. Tergites 1 silvery pollinose with 10–12 dark bristles. Abdomen dissected. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE:MWE = 1.4). Surstyli widened at the base, elongated, moderately narrowed before apex, pointed and bent to each other at the apex ( Fig. 80A ). Genital capsule in ventral view: both gonopods equal in height, extended towards surstyli; phallic guide with 2–4 differently sized spines dorsolaterally; subepandrial sclerite long ( Fig. 80B ); Genital capsule in lateral view: both surstyli widened at base, moderately curved towards sternite ( Fig. 80D–E ); ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 80C ). FEMALE: unknown.

Etymology: The specific name is derived from the Latin word ‘prolate’, meaning elongated, referring to the long surstyli.

Examined material: HOLOTYPE: AUSTRALIA: South Australia: Flinder’s Ranges National Park , Brachina Gorge, Heysen Hilltop , 31°20’S, 138°33’E, 9.X.1997, J.&A. Skevington & C. Lambkin, JSS8392 (1♂, QM) ; PARATYPES: AUSTRALIA: Queensland: Culgoa Floodplain National Park , 7km WNW Toulby Gate ( TOLB6 M), 28°57’S, 146°52’E, Callitris , 19.III–20.V .2010 , C. Kelly & A. Coward, Malaise trap, 19280, CNC575034 (1♂, QM); Mount Moffatt summit, 25°4’S, 148°3’E, 18.XI.2014, J.H., A.M. & A.W. Skevington, CNC374933 (1♂, CNC); Brisbane, Mount Coot-tha , 27°29’S, 152°57’E ,, 250m, hilltop , 21.VIII .2004 , J. Skevington, hand collected, JSM811–3; JSS15388–90 (3♂, CNC).

Distribution: Australia (Queensland, South Australia) ( Fig. 135 ).

Notes: All but one of the specimens was captured hilltopping.

Figures

FIGURE 80. Male genitalia of Tomosvaryella prolata sp. nov. (CNC374933, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view.

FIGURE 135. Distribution of Tomosvaryella prolata sp. nov., T. pseudophanes, T. setaria sp. nov. in Australia.