Tomosvaryella prolixa Motamedinia, Skevington & Földvari, 2023

Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, Zootaxa 5599 (1), pp. 1-271 : 179-180

Tomosvaryella prolixa Motamedinia, Skevington & Földvari sp. nov.

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Figs 81A–E , 134

Diagnosis: This species can be recognized by the shape of the surstyli, widened in the basal half and narrowed and bent towards the sternites in the middle in dorsal view ( Fig. 81A ); extremely elongated hypandrium; gonopods extended towards surstyli, rounded at apex in ventral view ( Fig. 81B ).

Description: MALE: Body length: 2.5 mm. Head. Flagellum tapering; yellow-brown pollinose, scape and pedicel brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.1 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1/3, covered by scattered short dark bristles. Thorax. Postpronotum pale, with 2 long dark bristles. Mesonotum (viewed obliquely from front) brown, silvery pollinose with some long supra-alar bristles behind postpronotum lobes and two rows dorsocentral bristles. Scutellum brown, silvery pollinose with 4–6 long erect bristles (longer than dorsocentral bristles) along distal edge. Halter, knob, stem pale, base brown. Legs. Mid coxa with 3–4 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright yellow. Fore and mid trochanters with three and two erect bristles, respectively. Hind trochanter gray pollinose without distinct projection; two distinct ventrobasal bristles present on fore and mid femora. Hind femur without ventroapical spines, except 5–8 equally spaced brown bristles posteroventrally, longest up to 2/3 the width of hind tibia at distal end. Short distal spines present on first four tibiae. Hind tarsomeres not flattened, hind metatarsus almost as long as 2–4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1–2 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 grey-brown pollinose. Dispersed short bristles on all tergites present. Lateral bristles on first tergite present, 4–5 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: ST8 longer than wide, epandrium longer than wide (MLE:MWE = 1.2). Surstyli widened in basal half, strongly narrowed in apical half, right one narrower and longer in apical half, both surstyli bent towards sternite by 90° in apical half ( Fig. 81A ). Genital capsule in ventral view: both gonopods extended towards surstyli, rounded at apex, hypandrium extremely elongated; phallic guide strong and rather wide, with one small dorsal spine before apex ( Fig. 81B ); Genital capsule in lateral view: both surstyli straight and wide in basal half, then narrowed and bent towards sternite in the middle ( Fig. 81D–E ); ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 81C ).

FEMALE: Unknown.

Etymology: From the Latin prolixus, stretched out long, in reference to the elongate hypandrium of this species.

Examined material: Holotype: AUSTRALIA: New South Wales: Ledknapper National Reserve, 7.3km ENE Beulah Headquarters (LDN2M), 29°21’S, 146°13’E, E. melanopholia , Spinifex , 10.XII.2009 – 18.III.2010, S. O’Sullivan & K. Taylor, Malaise trap, CNC576520 (1♂, QM).

Distribution: Australia (New South Wales) ( Fig. 134 ).

Figures

FIGURE 81. Male genitalia of Tomosvaryella prolixa sp. nov. (CNC576520, QM) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view.

FIGURE 134. Distribution of Tomosvaryella patula sp. nov., T. powerae sp. nov., T. prolixa sp. nov. in Australia.