Tomosvaryella miltocentra Motamedinia, Skevington & Földvari, 2023

Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, Zootaxa 5599 (1), pp. 1-271 : 145-146

Tomosvaryella miltocentra Motamedinia, Skevington & Földvari sp. nov.

urn:lsid:zoobank.org:act:69ECA24E-347E-43D4-AD5A-8772C37C4D3D

Figs 65A–E , 131 , 144D

Diagnosis: This species can be recognized by the hind trochanter having a keel covered by some short spines along its edge ( Fig. 144D ); mid metatarsus flattened at its base, extended inward; elongated surstyli, curved to each other from middle to apex in dorsal view ( Fig. 65A ); gonopods extended towards surstyli, subepandrial sclerite distinct with a fin-shaped projection in its center in ventral view ( Fig. 65B ); right surstylus with hump-like edge dorsomedially in lateral view ( Fig. 65D–E ).

Description: MALE: Body length: 3.2 mm. Head. Flagellum tapering, pale at the apex, light brown at the base; scape and pedicel dark. Pedicel with two short dark upper bristles. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1/3. Thorax. Postpronotum pale, with 1–2 short dark bristles. Mesonotum (viewed obliquely from front) brown pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Halter, knob pale, tip brownish, stem brown. Legs. Mid coxa with 1–2 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/6 of tibiae yellow (tibiae otherwise brown), tarsal segments brown. Hind trochanter with a keel covered by some short spines along its edge ( Fig. 144D ). Ventrobasal spines absent on fore and absent on mid femur. Hind femur without ventroapical spines, except 3–6 short bristles posteroventrally. Subapical (distal) spines on first four tibiae present (short). Mid metatarsus flattened at its base, extended inward. Hind tarsomeres not flattened, hind metatarsus almost as long as 2–4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.1 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1–2 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites black, tergite 1 silvery brown pollinose, others brown pollinose. Lateral bristles on first tergite present, 4–6 long pale bristles up to as long as of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE:MWE = 1.8). Surstyli elongated, widened at the base with two small lobes ventrobasally. curved to each other at apex, left surstylus is longer than right one ( Fig. 65A ). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, rounded at apex, gonopods with a small semicircular projection on inner margin; phallic guide unclear, perhaps detached; subepandrial sclerite distinct with a fin-shaped projection in its center ( Fig. 65B ); Genital capsule in lateral view: both surstyli curved towards sternite, right surstylus with hump-like edge dorsomedially ( Fig. 65D–E ). Ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 65C ).

FEMALE: Unknown.

Etymology: From the Greek miltos, red earth and kentron, centre, in reference to the Red Centre (Alice Springs Region) of Australia where this specimen was collected.

Examined material: HOLOTYPE: AUSTRALIA: Northern Territory: Waterhouse Range , 39 km SW by S of Alice Springs, 24°1’S, 133°25’E, 11.X.1978, D.H. Colless, JSS9239 (1♂, ANIC).

Distribution: Australia (Northern Territory) ( Fig. 131 ).

Figures

FIGURE 65. Male genitalia of Tomosvaryella miltocentra sp. nov. (JSS9239, ANIC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view.

FIGURE 131. Distribution of Tomosvaryella macrostyla sp. nov., T. merinator sp. nov., T. miltocentra sp. nov. in Australia.

FIGURE 144. Hind legs of Tomosvaryella species in lateral view A) T. danielsi sp. nov., paratype (JSS9708, QM), B) T. gaimarii sp. nov., holotype (JSS16633, QM), C) T. melaleuca sp. nov., holotype (CNC59696, QM), D) T. miltocentra sp. nov., holotype (JSS9239, ANIC), E) T. wyperfeldensis sp. nov., holotype (CNC373426, QM), F) T. bounites sp. nov., paratype (JSS2217, QM).