Arhaphes Candèze, 1860

Parekar, Harshad & Patwardhan, Amol, 2025, Annotated catalogue of the click-beetle subfamilies Hemiopinae, Hypnoidinae, Lissominae, Negastriinae, Tetralobinae, and Thylacosterninae (Coleoptera: Elateridae) of India, Zootaxa 5679 (2), pp. 151-203 : 168-170

Genus Arhaphes Candèze, 1860

Arhaphes bicoloris ( Vats & Chauhan, 1993) SDI

Senodonia bicoloris Vats & Chauhan, 1993: 39 (original description; published type locality: Boom , India).

Orientis bicoloris [sic] Vats & Chauhan, 1993: Cate, 2007: 208 (erroneously treated it as Orientis bicoloris ; listed as species incertae sedis; noted as “possibly Hemirhaphes [sic]”, unavailable genus name, incorrect subsequent spelling of Hemirrhaphes not in prevailing usage- ICZN 1999:Art. 33.3; catalogue; “UP” as India: Uttaranchal, Uttar Pradesh); Gupta et al., 2018: 503 (catalogue; “2B” as India: Himachal Pradesh, Uttarakhand).

Arhaphes bicoloris ( Vats & Chauhan, 1993) : Kundrata et al., 2018b: 282 (transferred from Senodonia ; catalogue; India: “ Uttarakhand ”).

Type Locality. India: Boom, Pithoragarh District , Uttarakhand [Uttar Pradesh: Boom] .

Type Depository. Holotype, SU (??).

Distribution in India. Uttarakhand (Pithoragarh District).

General Distribution. India.

Remark. As the presence of this species in Sikkim and Uttar Pradesh requires confirmation, these localities are not included here. The species is considered endemic to India.

Arhaphes lineicollis Fleutiaux, 1933 SDI

Arhaphes lineicollis Fleutiaux, 1933: 10 (original description; published type locality: Coorg, India); Chatterjee, 1934: 10 (distributional record; India: Karnataka).

Type Locality. India: Kushalnagara, Kodagu District , Karnataka [Fraserpet, Coorg] .

Type Depository. Type, SU (??).

Material examined. 15 specimens were examined from Kodagu District (same locality as published), Karnataka ( FRID) .

Distribution in India. Karnataka (Kodagu District).

General Distribution. India.

Remark. The type material may be housed in MNHN, but verification was not possible. Fleutiaux (1933: 10) examined a total of 20 specimens when establishing this species, 15 of which are currently in FRID. As none of these specimens are marked as type, we have refrained from designating them as syntypes or a lectotype. The species is considered endemic to India.

Arhaphes luteipes Candèze, 1897 SDI

Arrhaphes luteipes Candèze, 1897: 54 (unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage- ICZN 1999: Art. 33.3; original description; published type locality: Mysore, India); Schwarz, 1906: 159 (unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage— ICZN 1999: Art. 33.3; catalogue; India: “Mysore”); Schenkling, 1925: 218 (unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage- ICZN 1999: Art. 33.3; catalogue; India: “Mysore”).

Arhaphes luteipes Candèze, 1897 : Stibick, 1971: 390 (corrected genus spelling); Schimmel & Tarnawski, 2012a: 540, 541 (incorrect application of ICZN 1999: Art. 51.3; incorrectly used parenthesis for author; “Mysore”).

Type Locality. India: Mysuru District [Mysore], Karnataka .

Type Depository. 1 Syntype, SU ( RBINS) .

Material examined. One specimen was examined from Karavali Stretch , Karnataka ( BNHS) ; one specimen was examined from Palakkad District , Kerala ( KFRI) ; seven specimens ( Fig. 3 ) were examined from Coimbatore District , Tamil Nadu ( FRID) .

Distribution in India. Karnataka (Karavali Stretch, Mysuru District), Kerala (Palakkad District), Tamil Nadu (Coimbatore District).

General Distribution. India, Thailand ( Schimmel & Tarnawski 2012a).

Remark. This represents the first published record of this species from Kerala & Tamil Nadu.

Arhaphes minusculus Candèze, 1878 SRI

Arrhaphes minusculus Candèze, 1878 : cxlii (unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage— ICZN 1999: Art. 33.3; original description; published type locality: Himalaya).

Arhaphes minusculus ( Candèze, 1878) : Schimmel & Tarnawski, 2012a: 539, 541 (incorrect application of ICZN 1999: Art. 51.3; distributional record; India: Sikkim, West Bengal).

Type Locality. Himalaya*.

Type Depository. Type, SU (??).

Distribution in India. West Bengal (Darjeeling District).

General Distribution. India, Laos, Thailand ( Schimmel & Tarnawski 2012a).

Remark. The type material is not in RBINS (pers. comm. J. Constant, RBINS). Schimmel & Tarnawski (2012a: 539, 541) proposed a new combination using the correct genus name. According to ICZN 1999: Art. 51.3.1, parentheses are not to be used if the species-group name was originally combined with an incorrect generic name. As the presence of this species in Sikkim requires confirmation, this locality is not included here.

Arhaphes opacus Candèze, 1878 SRI

Arrhaphes opacus Candèze, 1878 : cxlii (unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage- ICZN 1999: Art. 33.3; original description; published type locality: Himalaya); Candèze, 1891: 120 (catalogue; “Himalaya”); Candèze, 1892: 491 (unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage- ICZN 1999: Art. 33.3; distributional record; India: Jharkhand); Schwarz, 1906: 159 (unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage- ICZN 1999: Art. 33.3; catalogue; “Himalaya”); Schenkling, 1925: 218 (unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage- ICZN 1999: Art. 33.3; catalogue; “Himalaya”).

Hemirrhaphes opacus ( Candèze, 1878) : Ôhira & Becker, 1973: 69 (transferred to Hemirrhaphes ).

Arhaphes opacus ( Candèze, 1878) : Schimmel & Tarnawski, 2012a: 539 (incorrect application of ICZN 1999: Art. 51.3; distributional record; India: Sikkim, West Bengal).

Type Locality. Himalaya*.

Type Depository. Type, SU (??).

Distribution in India. Jharkhand (Gumla District), Sikkim (Gyalshing District), West Bengal (Darjeeling District).

General Distribution. India, Nepal ( Ôhira & Becker 1973; Cate 2007).

Remark. The type material may be housed in RBINS, but verification was not possible. Schimmel & Tarnawski (2012a: 539) proposed a new combination using the correct genus name. According to ICZN 1999: Art. 51.3.1, parentheses are not to be used if the species-group name was originally combined with an incorrect generic name. Based on the information provided by Candèze (1892: 481) and Hayek (1973: 273), the locality is added as Jharkhand.

Arhaphes ruficollis Candèze, 1892 SDI

Hemirrhaphes ruficollis Candèze, 1892: 491 (original description; published type locality: Barwai , India).

Arrhaphes ruficollis ( Candèze, 1892) : Schwarz, 1906: 159 (transferred to Arrhaphes , unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage- ICZN 1999: Art. 33.3; catalogue; India: “Chota-Nagpore”); Schenkling, 1925: 218 (unavailable genus name, incorrect subsequent spelling of Arhaphes not in prevailing usage- ICZN 1999: Art. 33.3; catalogue; India: “Chota Nagpore”).

Arhaphes ruficollis ( Candèze, 1892) : Schimmel & Tarnawski, 2012a: 539 (incorrect application of ICZN 1999: Art. 51.3; distributional record; India: Meghalaya).

Type Locality. India: Barwai , Jharkhand .

Type Depository. Type, SU (??).

Distribution in India. Jharkhand (Gumla District), Meghalaya (West Garo Hills District).

General Distribution. India.

Remark. The type material may be housed in RBINS, but verification was not possible. Schimmel & Tarnawski (2012a: 539) proposed a new combination using the correct genus name. According to ICZN 1999: Art. 51.3.1, parentheses are not to be used if the species-group name was originally combined with an incorrect generic name. Based on the information provided by Candèze (1892: 481) and Hayek (1973: 273), the locality is added as Jharkhand. The species is considered endemic to India.

Figures

FIGURES 1–6. Elateridae from India, dorsal view. 1, Hypnoidus beckeri Stibick, 1980; 2, Lissomus mastrucatus Gerstaecker, 1860; 3, Arhaphes luteipes Candèze, 1897; 4, Miquasus improvisus (Dolin, 2001); 5, Zorochros mussardi Dolin & Cate, 1998; 6, Cussolenis wallacei Muona, 2020.