Ceutorhynchus cochleariae (Gyllenhal, 1813)

Korotyaev, B. A., 2020, A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae), Zoosystematica Rossica (China) 29 (2), pp. 353-367 : 359-360

Ceutorhynchus cochleariae (Gyllenhal, 1813)

( Figs 8, 10, 23)

Distribution. Sub-boreal Western Palaearctic species distributed from West Europe to Irkutsk Prov. ( Korotyaev, 1980) and northern Tuva ( Korotyaev, 2019). Dieckmann (1972) and Colonnelli (2004) report it from Anatolia (Asian part of Turkey) but I have not seen material from Turkey. The record from Georgia in Colonnelli (2004) is based apparently on the only known specimen from the City of Poti on the Black Sea coast in Western Georgia ( Korotyaev & Cholokava, 1989; see taxonomic notes below).

Hosts. Usually Cardamine pratensis ( Dieckmann, 1972: 70) , but species of several other genera of Brassicaceae were also listed in the cited publication by Dieckman of which E. Colonnelli mentions only Dentaria enneaphyllos and Lunaria rediviva ( Colonnelli, 2004) . In Tuva the species was found only on Cardamine pratensis out of the 28 investigated in 1979 and 1980 species of crucifers ( Korotyaev, 2019). In Irkutsk and its vicinities ( Korotyaev, 2019), I have collected beetles from Draba nemorosa and Barbarea arquata far from water bodies in the habitats where no Cardamine sp. was present.

Taxonomic notes. The taxonomy of C. cochleariae in the Caucasus needs investigation. This species is apparently absent from the largest part of the steppe zone of European Russia ( Arzanov, 2015), but a slightly different form with paler (reddish) legs and denser and finer dorsal vestiture is similar to C. cochleariae but with darker legs with black femora and tibiae and rather bright reddish tarsi; it also has sparser dorsal vestiture, and the beetles look darker than northern C. cochleariae . So, three allopatric forms exist along the northsouth transect from the forest zone of European Russia through the Caucasus foothills to the middle forest belt with Picea orientalis and Abies nordmanniana .

common in April and early May on Cardamine spp. in the Kuban River flood plain in the Republic of Adygea south of Krasnodar City. The single specimen reported by Korotyaev & Cholokava (1989) from Poti is apparently conspecific with this form. In the piedmont and low-hill Northwestern Caucasus this form has not been collected for many years, and only C. filirostris (Reitter, 1888) occurs (usually in small numbers) on Cardamine spp. and Dentaria spp. , but more often on Alliaria petiolata . At higher elevations, usually at about 1300 m a.s.l. and above, C. filirostris is replaced (on Cardamine lazica in Karachai-Cherkess Republic: Korotyaev & Cholokava, 1989) by a form very