Physogaleus latus ( Storms, 1894 )

Szabó, Márton & Kocsis, László, 2016, A preliminary report on the Early Oligocene (Rupelian, Kiscellian) selachians from the Kiscell Formation (Buda Mts, Hungary), with the re-discovery of Wilhelm Weiler’s shark teeth, Fragmenta Palaeontologica Hungarica 33, pp. 31-64 : 54-56

Physogaleus latus ( Storms, 1894)

(Figs 69–73)

1894 Protogaleus latus n. sp. – STORMS, p. 78, pl. 6, figs 17a-c.

1938 Eugaleus latus Ler. – WEILER, p. 8, pl. 1, figs 10, 11.

1938? Physodon contortus G. var. hassiae Jkl. – WEILER, p. 8, pl. 1, figs 19, 20.

1996 Physogaleus latus ( Storms, 1894) – MÜLLER, pp. 39–40, pl. 1, figs 3a-c, 7a-b.

1999 Physogaleus latus ( Storms, 1894) – MÜLLER, pp. 52–53, pl. 6, figs 1–4.

2001 Physogaleus latus ( Storms, 1894) – REINECKE et al., pp. 30–32, pls 46–49.

2010 Physogaleus latus (Storms, 1874) – HOVESTADT et al., p. 60, figs 15–28.

2014 Physogaleus latus ( Storms, 1894) – REINECKE et al., pp. 34–35, pls 31–34.

Referred material: 3 teeth (V.61.761., VER 2016.3435., VER 2016.3448.)

Remarks: The dentition of Physogaleus has dignathic and gradient monognathic heterodonty ( REINECKE et al. 2014). The anteriors are nearly as high as wide, while the lateral-distal teeth are wider than high. The mesial cutting edge is often serrated basally, and the serration vanishes from the half of the mesial cutting edge to the tip. The distal enamel shoulder has stronger serrations. The upper anterolaterals have a convex mesial cutting edge, while that of the lower anterolateral teeth is straight or weakly concave. The root is wide and low, it runs mesiodistally and bears a visual transversal groove. The species has been also reported by BAUT & GÉNAULT (1999), HOVESTADT & HOVESTADT-EULER (2010), HOVESTADT et al. (2010), and REINECKE et al. (2001, 2014).

Weiler also mentioned and figured another species, Physodon contortus G. var. hassiae Jkl. (1938, pl. 1, figs 19–20; also Fig. 69 of this work), which unfortunately was not found in the HNHM collection. Comparing this illustration with other faunas (e.g., REINECKE et al. 2014) it is highly possible that this specimen belongs to P.latus and represents an upper anterolateral tooth.

Another upper anterolateral tooth figured by WEILER (1938, pl. 1, fig. 10; also Fig. 71 of this work) has features typical for upper anterolaterals of the species. We suggest that specimen VER 2016.3448. (Fig. 70) is Weiler’s figured one, since it was found among other specimens of Weiler, and the preserved portions and size are the same. The major portion of the root and the serrated basal part of the mesial cutting edge are missing, but the first two serrations of the distal shoulder are preserved. The crown is wide, and shows a convex mesial cutting edge.

The lower anterolateral (V.61.761.; Figs 72–73) is sitting on a small piece of clay-matrix in labial aspect. A handwritten label (“ Eugaleus latus Ler. Taf. I Fig. 11 ”) was found under the specimen, which shows that this is one of Weiler’s figured ones (see WEILER 1933, pl. 1, fig. 11; also Fig. 73 of this work). Except some cracks, this tooth is in nearly perfect condition.

Figures

Figs 10–13. Notorynchus primigenius (Agassiz, 1843). – 10. Lower symphyseal tooth (VER 2016.3455.), lingual view. – 11. Specimen VER 2016.3455. as figured by WEILER (1938). – 12. Lower lateral tooth (V.61.672C.), lingual view.– 13. Suggested specimen V.61.672C., as figured by WEILER (1933).– Figs 14–17. Hexanchus agassizi Cappetta, 1976. – 14. Upper anterolateral tooth (V.61.285.), lingual view. – 15. Lower lateral tooth (V.61.770.), lingual view. – 16. Lower lateral tooth (VER 2016.3456.), labial view. – 17. Lower lateral tooth (VER 2016.3457.), lingual view. Scale bars: 5 mm