Trichotichnus Morawitz, 1863

Kataev, B. M., 2020, Description of a new subgenus of Trichotichnus (Coleoptera: Carabidae), with two new species from the Western Ghats (India), and remarks on other subgenera, Zoosystematica Rossica (China) 29 (2), pp. 172-194 : 190

Subgenus Trichotichnus Morawitz, 1863

Trichotichnus Morawitz, 1863: 63 [as genus; nomen protectum, see Bousquet (2008: 328)]. Type species: Trichotichnus longitarsis Morawitz, 1863 , by monotypy.

Asmerinx Tschitschérine, 1898: 183 [as genus]. Type species: Carabus laevicollis Duftschmid, 1812 , designated by Tschitschérine (1900).

Pteropalus Casey, 1914: 64 [as genus]. Type species: Harpalus vulpeculus Say, 1823 , designated by Habu (1954).

Velimius Jedlička, 1952: 51 [as genus]. Type species: Velimius edai Jedlička, 1953 , by monotypy.

Diagnosis. Fronto-ocular furrows shallow, at most deepened only at clypeus, reaching or not reaching supraorbital furrows. Fronto-clypeal suture superficial or slightly deepened. Genae narrow to moderately wide. Ligular sclerite widened at apex (Figs 39–41) or (for example, in Europe- an species) almost parallel-sided (Fig. 42). Paraglossa narrow, separated from ligular sclerite by wide (about as wide as paraglossa apically) notch. Elytra with lateral groove flat throughout, without distinct elongate convexity along this groove apically. Elytral marginal umbilicate series more or less continuous or with a short gap at middle. Metepisternum markedly narrowed posteriorly, its anterior margin not longer than inner margin. Abdominal sternite VII of male with one or two pairs of marginal setigerous pores. Gonocoxite with one short seta (occasionally with two setae) on ventral outer edge or on both ventral and dorsal outer edges at their middle or in their basal portion. Median lobe of aedeagus with apical orifice in dorsal position or slightly shifted to left; apical capitulum absent or small, in many species prominent only dorsally.

Composition. This subgenus comprises more than one hundred described species ranged mostly over Palaearctic East Asia and northern part of the Oriental region from northern Pakistan and northern India to the Russian Far East, Japan and Taiwan; south to Vietnam, Thailand and Laos; one species was described from the Philippines and one from North Borneo; four species occur in Central Europe and two in the eastern areas of North America. More than half of all known species were described from Japan; most of them belong to the brachypterous leptopus group.