Trichotichnus, Morawitz, 1863

Kataev, B. M., 2020, Description of a new subgenus of Trichotichnus (Coleoptera: Carabidae), with two new species from the Western Ghats (India), and remarks on other subgenera, Zoosystematica Rossica (China) 29 (2), pp. 172-194 : 191-192

Key to subgenera of Trichotichnus

1. Fronto-ocular furrows deep throughout, reaching inner margin of eyes and continuing into deep supraorbital furrows. Fronto-clypeal suture deepened, fused laterally with fronto-ocular furrows. Ligular sclerite narrow, not widened at apex, more or less parallel-sided or moderately narrowed apically. Paraglossa moderately wide or somewhat narrow, rounded apically, separated from ligular sclerite by narrow notch............. Bottchrus

– Fronto-ocular furrows shallow, at most deepened only at clypeus, reaching or not reaching supraorbital furrows, occasionally only slightly visible. Fronto-clypeal suture superficial or slightly deepened. Ligular sclerite narrow or widened at apex. Paraglossa wide or rather narrow, separated from ligular sclerite by narrow or wide notch........ 2

2. Paraglossa narrow (Figs 39–42), notch between it and ligular sclerite wide, about as wide as paraglossa apically. Ligular sclerite widened at apex or almost parallel-sided. Last visible abdominal sternite (VII) of male with one or two pairs of marginal setigerous pores.............................. 3

– Paraglossa wider ( Figs 4, 10, 18, 24, 31), notch between it and ligular sclerite narrow, much narrower than paraglossa apically. Ligular sclerite not widened apically, at most with outer angles slightly protruded laterally. Last visible abdominal sternite of male with two pairs of marginal setigerous pores....................................... 4

3. Metepisternum slightly narrowed posteriorly, its anterior margin longer than inner margin. Species from mainland China............ Amaroschesis

– Metepisternum markedly narrowed posteriorly, its anterior margin not longer than inner margin. Species from China and other countries........................................ Trichotichnus s. str.

4. Ligular sclerite ( Figs 18, 24, 31) parallel-sided or slightly narrowed before apex. Elytral marginal umbilicate series with more or less wide gap at middle, occasionally with one intermediate pore in gap, consisting of 15–20 setigerous pores. Body wider and flatter........................... Iridessus

– Ligular sclerite ( Figs 4, 10) markedly narrowed to apex. Elytral marginal umbilicate series continuous, consisting of 24–34 setigerous pores. Body more elongate and more convex.................................... Parairidessus subgen. nov.