Bottchrus Jedlička, 1935

Kataev, B. M., 2020, Description of a new subgenus of Trichotichnus (Coleoptera: Carabidae), with two new species from the Western Ghats (India), and remarks on other subgenera, Zoosystematica Rossica (China) 29 (2), pp. 172-194 : 191

Subgenus Bottchrus Jedlička, 1935

Bottchrus Jedlička, 1935: 8 [as genus]. Type species: Bottchrus philippinus Jedlička, 1935 , by monotypy.

Bellogenus Clarke, 1971: 264 [as genus]. Type species: Bellogenus amazeus Clarke, 1971 , by original designation.

Pseudotrichotichnus Habu, 1973: 225 [as a subgenus of Trichotichnus ]. Type species: Trichotichnus nanus Habu, 1954 , by original designation.

Diagnosis. Fronto-ocular furrows deep throughout, reaching inner margin of eyes and continuing into deep supraorbital furrows. Fronto-clypeal suture deepened, fused laterally with fronto-ocular furrows. Genae moderately wide. Ligular sclerite narrow, not widened at apex, more or less parallel-sided or moderately narrowed apically. Paraglossa moderately wide or somewhat narrow, rounded apically, separated from ligular sclerite by narrow notch. Elytra with lateral groove flat throughout or (in some species), with elongate convexity along this groove apically forming short additional interval there. Elytral marginal umbilicate series with more or less wide gap at middle. Abdominal sternites without extra setae or with very fine and short setae medially. Abdominal sternite VII in male and female with two pairs of marginal setigerous pores. Gonocoxite with one or two stout setae on both ventral and dorsal outer edges at middle or basally. Median lobe of aedeagus with apical orifice shifted to left and with apical capitulum in most species.

Composition. This subgenus includes more than 70 described species. Most of them are distribut- ed over Southeastern Asia from Sri Lanka and Hindustan to southern China and Japan, south to Australia. One species, T. amazeus ( Clarke, 1971) , is known from Ethiopia. The systematic position of some species described from New Guinea and Australia needs further study.

Remarks. Morphologically well defined subgenus which is easily distinguished from other subgenera by having fronto-ocular furrows deep throughout, reaching deep furrows around inner margin of eyes ( Habu, 1973; Noonan, 1985; Kataev, 2016).