Trigonella bakhtiarica Ranjbar & Z. Hajmoradi., 2015

Ranjbar, Massoud & Hajmoradi, Zahra, 2015, A new species of Trigonella sect. Ellipticae (Leguminosae-Papilionoideae) from Iran, including cytogenetic and anatomical notes, Phytotaxa 202 (1), pp. 26-34 : 31-32

Trigonella bakhtiarica Ranjbar & Z. Hajmoradi. sp. nov. Figs. 1 , 2 .

Affinis Trigonella aphanoneura planta 38–58 cm (nec ad 30 cm) alta, vexillum 9–10 mm (nec ± 7 mm) longum, leguminibus ovato lanceolatis (nec oblongis), 7–12 mm (nec 20–25 mm) longis differt.

Type:— IRAN. Prov. Chahar Mahal Va Bakhtiar, between Naghan and Boroujen, Sabz-kuh protected area, Chahartagh , 31.70°N, 50.80°E, 2200 m, 5 June 2012, Ranjbar 33055, (holotype BASU, photo W ).

Perennial, 38–58 cm tall. Stem branched, glabrous or occasionally sparsely covered with appressed white hairs 0.2–0.5 mm long. Stipules herbaceous, 2–5 × 0.6–1 mm, adnate to the petiole for up to 0.5 mm, lanceolate, glabrous. Leaves 3-foliolate, 8–17 mm long; petiole 2–5 mm long, glabrous to sparsely appressed hairy, similar to the stem. Leaflets obdeltoid to obcordate, 4–10 × 3–7 mm, truncate or emarginate, glabrous above, glabrous or sparsely appressed white hairy beneath, the hairs 0.2–0.5 mm long. Inflorescence an erect raceme, laxly 3–6-flowered, apex aristate; peduncles 1.3–5.2 cm long. Pedicels 3–5 mm long. Bracts hyaline-membranous, 0.3–0.4 × ca. 0.2 mm, triangular, glabrous. Calyx ca. 5 mm long, tubular, glabrous or rarely with sparse appressed hairs 0.2–0.5 mm long; teeth filiform to subulate, ca. 2 mm long. Corolla yellow. Standard 9–10 × 5–6 mm, obovate, tapering abruptly into the short cuneate claw, apex rounded. Wings ca. 10 mm long; blades oblong, apex rounded, 6–7 × ca. 1.5 mm, claw linear, 0.6–0.7 mm long. Keel 8–9 mm long; blades 5 × 2.5 mm; claw linear, 3–4 mm long. Stamens diadelphous (9 fused + 1 free), 8–9 mm long. Ovary 5–6 mm with stipe ca. 1 mm long, glabrous. Pods 7–12 × 3–4 mm, ovate to lanceolate, widely rounded at the lateral side, with a slender stipe 1 mm long, at the apex obtuse to rounded, beaked; valves straw-colored to pale brown to cream, rugose with shallow ridges and furrows, glabrous. Seeds 1–2, oblong, ca. 3 × ca. 2 mm, greenish to brown.

Phenology: —Collected in flower and fruit in May and June.

Etymology: —The species is named after the type locality, Bakhtiari, Iran.

Distribution and habitat: — T. bakhtiarica is a narrowly distributed endemic species, known only from the type locality (fig. 2). It was collected from a stony area in the mountainous region of the Sabz-kuh protected area. T. aphanoneura , the closest relative of T. bakhtiarica , is common in the dry-steppe of Boroujen protected area and a few localities in the Esfahan Province.

Figures

FIGURE 1. Holotype of Trigonella bakhtiarica Ranjbar & Hajmoradi (Ranjbar 33055, BASU). A. habit with fruit. B. calyx. C. standard. D. wings. E. keel. F. ovary. G. androecium. H. pod of T. bakhtiarica. I. pod of T. aphanoneura. J. mitotic cell in T. bakhtiarica with 16 chromosome (2n = 2x = 16). K. meiotic cell in T. aphanoneura with 8 bivalents chromosome (2n = 2x = 16). Scale A = 2 cm, B–G = 1 cm, H–I = 5 mm, J–K = 2 μm

FIGURE 2. Distribution of Trigonella bakhtiarica (●) and T. aphanoneura (■) in Iran.