Bonea Roewer, 1914

Kury, Adriano B. & Machado, Glauco, 2025, Redescription of Bonea sarasinorum from Sulawesi, and the relationships of Bonea within Podoctinae (Opiliones, Grassatores, Podoctidae), Zootaxa 5659 (1), pp. 133-145 : 135-136

Bonea Roewer, 1914

• Bonea Roewer 1914: 87 ; Roewer 1923: 160; Suzuki 1982: 205; Zhang et al. 2013: 204 .

[Type-species by monotypy: Bonea sarasinorum Roewer, 1913 .]

• Zmissolus Roewer 1927: 311 . [Junior subjective synonym of Bonea Roewer, 1914 by Suzuki (1977: 37). Type-species by monotypy: Zmissolus cippatus Roewer, 1927 .]

• Posisus Roewer 1949: 258 . [Junior subjective synonym of Bonea Roewer, 1914 by Suzuki (1977: 37). Type-species by original designation: Posisus albertus Roewer, 1949 .]

• Kappacola Roewer 1949: 260 . [Junior subjective synonym of Bonea Roewer, 1914 by Suzuki (1977: 37). Type-species by original designation: Kappacola silvestris Roewer, 1949 .]

• Suraplus Roewer 1949: 262 . [Junior subjective synonym of Bonea Roewer, 1914 by Suzuki (1977: 37). Type-species by original designation: Suraplus palpalis Roewer, 1949 .]

• Parabonea Roewer 1949: 262 . [Junior subjective synonym of Bonea Roewer, 1914 by Suzuki (1977: 37). Type-species by original designation: Parabonea scopulata Roewer, 1949 .]

• Nurullus Roewer 1949: 262 . [Junior subjective synonym of Bonea Roewer, 1914 by Suzuki (1977: 37). Type-species by original designation: Nurullus armatissimus Roewer, 1949 .]

Composition. The genus comprises 10 species ( Kury et al. 2024) distributed across Borneo, Hainan Island, Java, the Malay Peninsula, the Philippines, and Sulawesi. However, based on the current diagnosis, not all these species should be included in Bonea , highlighting the need for a taxonomic revision.

Emended diagnosis

This diagnosis is based on the type species of Bonea , because the pertinence of the others still needs to be assessed on a case-by-case basis. Bonea sarasinorum belongs to the clade Podoctinae + Erecananinae based on: (1) the presence of a tower-like interocular mound, contrasting with Ibaloniinae , which typically lack such a feature, having either no interocular structure, only an interocular spine, or a less pronounced mound; and (2) femur I being S-shaped and armed with both a dorsal and a ventral row of numerous strong megaspines, whereas in Ibaloniinae , there are only a few ventral megaspines. Within the clade Podoctinae + Erecananinae , B. sarasinorum is more similar to Dongmoa and Erecanana based on: (1) the outline of the dorsal scutum in lateral view being thicker anteriorly (as opposed to elliptical in Podoctis , Baramia , and Hoplodino , or flattened in Metapodoctis and the reclinobunoids sensu Kury & Machado 2018); and (2) the presence of cervical tubercular bridges (CEtb) connecting the tower to the posterior part of the carapace. Bonea sarasinorum is more similar to Dongmoa specifically due to the presence of TS1. Bonea sarasinorum differs from Dongmoa and Erecanana in the following ways: (1) the presence of an elongated rectangular cleft at the apical border of the truncus penis, splitting it into two valves; and (2) a very small proximal capsula interna, buried between the valves.