Protadelomys cf. alsaticus, Hartenberger, 1969
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publication ID |
https://doi.org/10.1186/s13358-022-00245-3 |
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persistent identifier |
https://treatment.plazi.org/id/497F1B1D-FF8C-DF7E-C483-4546FB64FD19 |
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treatment provided by |
Felipe |
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scientific name |
Protadelomys cf. alsaticus |
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? Protadelomys cf. alsaticus morphotype 3
Material and measurements. (Additional file 1: S4; Table 4B). We refer two DP4, one P4, one M2, one M3; one m1, two m2, and three m3 to this morphotype.
Description
Upper teeth
DP4. Te larger DP4 (CSBX 2: ( Fig. 40a View Fig ).) of the Bouxwiller assemblage displays a more intricate pattern than the other DP4 ( Fig. 40b View Fig ). Te paraconule is relatively reduced and aligned with the straight protoloph, while the metaconule is duplicated, as are the mesostyle and the mesoloph. Te metalophule I joins the mediobuccal ridge of the metaconule, together with the main mesoloph. Te outskirts of the crown display irregular wrinkles, thicker than the thin ornamentation of? P. alsaticus .
P4. ( Fig. 40c, d View Fig ). CSBX 12 and 13 show common features with the P4 of?P.. alsaticus , but still differs from them in some aspects. Tey show a longer anteroloph with a preparacrista, which is stronger; the connection between the postparacrista, the small mesostyle, and the long premetacrista limits the shallow mesoflexus buccally. Te metalophule I is reduced or almost absent. Te hypocone is almost as large as the protocone.
M2. ( Fig. 40e View Fig ). BUX 67–3 is weakly worn, especially in its medial part. Te extra-ridges are more numerous than on the M1 BUX 67–35 (morphotype 1). Te paracone is bulged and the postparacrista is present. Te protoloph turns a little distally, buccal to the paraconule. Te latter separates from the protoloph by an oblique notch; it is stretched forward, inside the anteroflexus. Te paraconule attaches to the short transverse lingual part of the protoloph, linked at mid-protocone, just mesially to a low protocrista. Te premetacrista, the metacone, and the metalophule II merge into a loph, which is arcuate up to the metaconule level. Te metaconule is complex, which makes it difficult to identify a swollen conule present in this area alongside at least three very close ridges. Te mesostyle is complex, prolonged by a bifurcated mesoloph. A ridge extends from the end of the buccomesial border of the metacone and runs parallel to the mesoloph. Another ridge descends from the metalophule II in the basin; it could constitute a remnant of the metalophule I. Te endoloph is high and thick. Te hypocone is more buccal than the protocone. It separates from a distinct posteroconule by a shallow and narrow slot. Te posteroloph is very short.
M3. ( Fig. 40f View Fig ). Te unworn M3 BUX 67–27 appears more concave than the other molars, as the cusps and the surrounding bucco-distal ridge or the lingual one are not high or thick. Te paracone is not much prominent, and only slightly higher than the protoloph. Te mesially protruding slender paraconule is oriented towards the anterostyle. Te latter, not well individualized, only represents a swelling at the extremity of the preprotocrista. Te extra-ridges and the granules are numerous on the mesoflexus floor. Te metacone is low, not salient, and located above the bucco-distal crest joining the mesostyle area. Tis metacone is similar in height as the posteroloph, which is short. Te protocone is a little elongated mesiodistally from the preprotocrista to the postprotocrista. A thickening, which actually corresponds to the high and strong endoloph observed on the M2, prolongs the postparacrista (BUX67-3). A linguobuccal lophid, as high as the paraconule, faces the protocone and reaches the central extra-ridges. Te hypocone is not clearly identifiable, as it is not swollen; it is strongly linked to the postprotocrista. Tere is no trace of a sinus. Te metaconule is lower than the paraconule, and reduces to a thin ridge postwardly oriented. Smooth vertical wrinkles, diverging from its occlusal surface, adorn the lingual surface of the protocone.
LoWer teeth
m1–m2. ( Fig. 40g, h, i View Fig ). Te m1 BW 56–59 has numerous extra-ridges and wrinkles. Te elements of the postprotocristid + ectolophid area are in line obliquely, mesiobuccal to distolingual, like on the two m2 (BUX 67–129 and 142). BUX 67–129 is little worn, as are their short anteroconid whereas the premetacristid (mesial lingual metalophulid I) is longer than on the m1 and it bears two long mesiodistal ridges, parallel to the one starting at the lingual end of the buccal metalophulid I. Te postmetacristid is steep and long. On BUX 67–142
( Fig. 40i View Fig ), which is not too much worn but a little damaged, the premesoconid–mesoconid–postmesoconid elements are obliquely in line with the postprotocristid, like on the m1. Te ectomesolophid is faint. Te lingual metalophulid I, transverse, is almost in line with the buccal one, without any break. Even worn, it is possible to distinguish numerous low extra-ridges in the central basin and a few wrinkles descending from the postprotocristid and mesoconid area in the sinusid.
m3. ( Fig. 40j, k View Fig ). BUX 67–169 differs from the other large m3 BUX 68–23 (morphotype 1) in its mesial part that is relatively wider than the distal one. It also differs from it in the presence of a mesostylid at the extremity of the postmetacristid closing lingually the mesoflexid, and in the presence of an entolophid. A few wrinkles descend buccally from the postmetacristid and from the mesostylid; some granules are visible in the basin, even if the tooth is worn. Te lingual metalophulid I is mesial (premetacristid) and bears two long low mesiodistal ridges running in the basin, parallel to the short ridge emitted by the lingual end of the transverse buccal metalophulid I. A premesoconid spur is well distinct, directed lingually from the end of the postprotocristid. Te lophid developed from the postprotocristid to the prehypocristid is mesially oblique and then oriented mesiodistally, with the distal part of the ectolophid. Te ectomesolophid is absent (contra on BSK-A25 where it is well present). Te buccal part of the entolophid connects to the strong postmesoconid ridge and to the prehypocristid. Te lingual entolophid is duplicated and the distal one is slightly directed distally. While worn, the hypoconulid seems to be also duplicated, with a distomesial ridge running towards the buccal entolophid. Te extra-ridges and granules are numerous in the low basins of BSK-A25.
Remarks and comparisons
Te larger lower molars of? Protadelomys cf. alsaticus morphotype 3 are similar in size as the ones of Masillamys parvus fom Messel. Tey share the numerous extra-ridges and the organization of the lingual metalophulid I area. However, the oblique postprotocristid + ectolophid is longer, the sinusid is deeper linguobuccally. Moreover the trigonid of p 4 in this species is better developed than in p4 of Masillamys parvus , with a distinct protoconid, and the entolophid and extraridges are thicker in the species from Bouxwiller than in M. parvus .
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