Protadelomys cf. alsaticus, Hartenberger, 1969
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publication ID |
https://doi.org/10.1186/s13358-022-00245-3 |
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persistent identifier |
https://treatment.plazi.org/id/497F1B1D-FF8A-DF05-C739-43E6FA36FC79 |
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treatment provided by |
Felipe |
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scientific name |
Protadelomys cf. alsaticus |
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? Protadelomys cf. alsaticus morphotype 1
Material and measurements (Additional file 4: S4, Table 4B). We refer two P4, one M1, two M2, two dp4, one p4, two m1, two m2, one m3, to this morphotype.
Description
Upper teeth
P4. ( Fig. 38a View Fig ) One P4 (CSBX 8) displays a simpler occlusal morphology than other specimens, and cusps more bulbous. Te paracone and metacone are very close to one another, and the mesoflexus is narrow. Te mesostyle is small, adjacent to the metacone, and the mesoloph is absent. Another P4 (CSBX 29) could be close to CSBX 8, with its bulbous cusps and small hypocone, but it is too damaged, especially in the area between the paracone and metacone.
M1. ( Fig. 38b View Fig ) One tooth (CSBX 35) has been questionably refered to an M2 of? P. alsaticus (Hartenberger, 1969: pl.2, Fig. 7 View Fig ). Tis weakly worn M1 displays an anterosyncline wider than that of the other M1. Te parastyle is strong, slightly separated from the anteroloph, and weakly connected to a sloping preparacrista. Te anteroloph is weakly worn, whereas the parastyle is unworn. Te anterostyle is strong, connected to both the preprotocrista and the anteroloph. Te endoloph is short and thick, nearly as high as the lingual main cusps. Te hypocone is slightly smaller than the protocone, its anterior and posterior arms being short. Te parastyle presents one additional ridge connected to the anteroloph in the anterosyncline. Te paracone is slightly swollen. Te buccal part of the rectilinear protoloph is as high as the paracone, and its lingual part is lower. Te protoloph connects to the protruding paraconule and to the middle of the protocone (protocrista). Te paraconule is mesial to the protoloph and slightly less swollen than the metaconule. Te paracone is framed by the pre- and post-paracristae. Te latter is strong, and interrupted anterior to the mesostyle. Te mesostyle prolongs in a strong mesoloph, joining a low cusp (mediobuccal) against the metaconule. A short ectocingulum is present on the distal flank of the mesostyle. Te premetacrista is strong and the metacone a little thicker than the paracone. Te metalophule II is high and the metalophule I is low, descending mesially from the latter to the base of the metaconule. Te connection between the metalophule II and the metaconule is low as is the one between the latter and the mid-hypocone. Tere is a strong posteroconule, lowly attached to the metaconule and to the mesiodistal post-hypocrista. Te sinus forms only a very shallow depression below the endoloph. Some rare very low ridges, present on the floor of the flexi, connect the main structures.
(See figure on next page.)
Fig. 36 View Fig Schematic representations of the variations of the outer wall components of the lower molars of? Protadelomys alsaticus Hartenberger,
1969 from Bouxwiller (Bas-Rhin).Color caption: the colors represent different cuspids, cristids or lophids: 1: Brown= protoconid; 2: buccal part of the metalophulid I=pale violet; 3: postprotocristid =brown like the protoconid; 4: premesoconid swelling (spur or ridge)= brown, at the distal end of the postprotocristid; 5: mesial ectolophid =green; 6: mesoconid = blue; 7: ectomesolophid =blue; 8: postmesoconid ridge and swelling =ochre;9 distal ectolophid= Green; 10: prehypocristid= black; 11: hypoconid =black;12: entolophid=black; 13: poshypocristid= black
M2. ( Fig. 38c View Fig ) Among the largest M2, BUX67-37 and 67–77 are referred to this morphotype. Tey show distinct and swollen conules. Te metaconule is simple. If the thick buccal metalophule II is transverse on BUX 67–37, it thickens lingually and curves distally on BUX 67–77 to join the posteroconule. Te mesostyle and the buccal mesoloph are present but their width seems to vary: on BUX 67–37, it is thicker and longer and reaches the metaconule.
LoWer teeth
dp4. ( Fig. 38d, e View Fig ) Two dp4 differs from the others by having a transverse metalophulid I, as well as a short anterolophid closing a small anteroflexid. On BUX 67–96, the mesiodistal ridge is barely marked, as is the buccal half of the entolophid. Te lingual part of the latter is short and distally oriented towards the posterolophid. CH 25 is much worn with a damaged mesial flank, but the “trigonid” area is preserved and similar to that of BUX 67–96.
p4. ( Fig. 38f View Fig ) On CH 18, the metaconid is strong and prolonged buccally in a high and short anterolophid up to the preprotocristid. A short transverse metalophulid I runs parallel to the anterolophid, both being separated by a minute anteroflexid. Te short postmetacristid descends abruptly towards the lingual opening of the mesoflexid, mesially to the entoconid, which is massive. Tis lingual opening is V shaped and the vertical surface of enamel is concave at the junction of the lingual flanks of metaconid and entoconid. Tis is due to the bulging of the buccal flanks of the metaconid + postmetacristid, and the bulging of the buccal flanks of the entoconid. One thick mesiodistal ridge runs from the metaconid to the premesoconid spur; lingual to it, there is an additional parallel crest, more discontinuous. Tree short extraridges fill the space between the main ones. Te ectolophid makes an angle behind the mesoconid; the latter is slightly stretched mesiolingual to bucco-distal. A short and strong ectomesolophid curves and its buccal end connects a hypoconid spur, which is mesiobuccal. Te lophids, ridges, and conids appear thick when worn.
m1–2. Te peculiar shape of the buccal wall of the p4 and m3 is not so easy to distinguish on the m1–2: the recess between the bulging flanks of the metaconid and the entoconid is not strongly marked, and varies with wear. Among the largest m2, BUX 67–138, 139: Fig. 38i View Fig , 142, and CSBX 53 show these features; BUX 67–121 does not. We found this character on the m1 BUX 67–125 and 137. BUX 67–125 has a short and swollen anteroconid; the lingual metalophulid I is mesial then turns distally in a short ridge, which meets the distomesial hook located at the end of the transverse buccal metalophulid I. Te weak and low additional lingual metalophulid I makes a long mesiodistal ridge up to the center of the basin, where it meets a few low and slender extra-ridges. Another extra-ridge runs on the buccal flank of the metaconid lingually to this additional metalophulid I. A short premesoconid spur is present. Te mesoconid is a little stretched obliquely (mesiolingual to bucco-distal); it bears a barely marked ectomesolophid and a weak mesiobuccal spur of the hypoconid. Its distal end is linked to a thick postmesoconid. Te distal ectolophid is low and very short, and this is especially clear on unworn and weakly worn teeth. Te entolophid is almost straight and connected to the prehypocristid. Te post-hypocristid is short, the hypoconulid strong, and the posterolophid displays two successive bulges. Te occlusal face of BUX-137 is damaged and all occlusal structures are worn.
BUX 67–138 is a pristine m2, reduced to its enamel cap ( Fig. 38h View Fig ). At its lingual end, the stretched anteroconid connects with a thin discontinuous mesiodistal ridge running towards the center of the basin. Te lingual metalophulid I is mesial and short as a premetacristid. Te buccal metalophulid I is longer, being transversely oriented at its most buccal part then turning slightly backwards. An additional lingual metalophulid I is thin and low, like the extra-ridges located in the basin. Te posprotocristid ends in a premesoconid spur. It strongly connects to the mesoconid, which is stretched buccomesial to linguodistal. A strong ectomesolophid descends mesiodistally along the buccal flank of the mesoconid, while a thin mesiobuccal spur climbs on the hypoconid flank. Two postmesoconid spurs emerge from a unique point at the distal end of the mesoconid. Te distal ectolophid is low, and connects to the short prehypocristid. Te entolophid is almost straight and complete; it is linked to the extremity of the prehypocristid. Te hypoconulid swells at the extremity of the post-hypocristid, and the posterolophid is long, relative to the post-hypocristid length.
m3. ( Fig. 38j View Fig ) BUX 68–23 is the larger m3 of the theridomorph assemblage from Bouxwiller. Its morphology is relatively simple when compared to other m3, and the extra-ridges are very low and rare. Its metaconid and entoconid are more massive. Te lingual metalophulid I is mesial as it is the case on nearly all the m3 from Bouxwiller; it is short and its buccal end joins the anterolophid lingual end. Te additional more distal lingual metalophulid I is slender and low; it descends in the direction of the transverse buccal metalophulid, without connecting it. Tere is no mesostylid. Te postprotocristid is long and thick, and connects to the mesoconid by a low mesial part of the ectolophid. Te mesoconid is a little stretched mesiodistally, with two tiny mesial and distal bulges with short lingual spurs. A thick linguodistal short ridge starts in front of the distal bulge. We interpret this ridge as the postmesoconid ridge. Indeed, there is no trace of a lingual part of the entolophid. Te entoconid is free from the posterolophid. Te latter is very short, making a short and thick curve with the hycoconulid and the post-hypocristid. Te hypoconulid is unique and not swollen.
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