Euryobeidia languidata ( Walker, 1862 )

Liu, Bo, Stüning, Dieter & Han, Hongxiang, 2025, Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes, ZooKeys 1260, pp. 313-343 : 313-343

Euryobeidia languidata ( Walker, 1862)

Figs 2–8 , 38 , 39 , 50 , 56

Abraxas languidata Walker, 1862, List Specimens lepid. Insects Colln Br. Mus. 24: 1122. Holotype, Nepal. ( NHMUK)

Euryobeidia languidata : Wehrli 1939, in Seitz, Gross-Schmett. Erde 4 (Suppl.): 269. Unavailable.

Euryobeidia languidata : Fletcher 1979, The Generic Names of Moths of the World 3: 84; Stüning, D. 2000, Moths of Nepal, part 6: 110; Xiang et al. 2017, Zootaxa 4317 (2): 371.

Type material examined.

Holotype. Nepal • ♀; ‘ Nepal’ ; Hardwicke Bequest; NHMUK. (Abdomen lost; see Fig. 2 )

Additional material examined.

Nepal • 1 ♀; Kathmandu Valley, Godavari ; 1600–1800 m; 5 Jun. 1967; leg. Dierl-Schacht; ZSM. India • 1 ♂; Khasis; May 1896; Nat. Coll.; coll. Wehrli, ZFMK. China – Fujian Province • 1 ♂; Quanzhou City, Dehua County, Jiuxianshan ; 1200 m; 12 May 2024; C. L. Huang & Z. Peng leg.; GenBank no.: PQ 083540; gen. prep. no. CRICATAS 00283 ; CRICATAS – Taiwan • 1 ♂; Nantou, Sang-Gan nr. Pu-li ; 23 ° 58 ' N, 120 ° 55 ' E; Apr. – Jul. 2008; local coll.; ZFMK. Vietnam • 1 ♂; N. Vietnam, Vinhu, Tam Dao ; 900 m; 29 May 1997; B. Tanaka leg.; CKY. Japan – • 1 ♂, 1 ♀; ‘ Japon, Wileman , 1898 ’; male gen. prep. Wehrli no. 5525, female gen. prep. no. 2460 - DS; ZFMK – Tokyo • 1 ♂; Tokio ; 17 Jun. 1910; gen. prep. Wehrli. no. 5517; ZFMK • 1 ♂; Abiko, Chiba Pref. nr. Tokyo ; May 1914; ZFMK • 1 ♂; Chiyoda, Kokyo ; 21 May 1998; Y. Kishida leg.; gen. prep. no. RS 8877; CRS / NIAES (shown by Sato 2011, pl. 1-015 - 9) • 1 ♀; Chiyoda, Kokyo ; 11 Jun. 1998; Y. Kishida leg.; gen. prep. no. RS 8878; CRS / NIAES (shown by Sato 2011, pl. 1-015 - 10) – Niigata Prefecture • 1 ♀; Niigata City, Nishikan-ku, Iwamuro , Parking of Yahiko ; 23 Jul. 2007; R. Sato leg.; CRS / NIAES (shown by Sato 2011, pl. 1-015 - 11) .

Remarks.

Euryobeidia languidata obviously is extremely rare at its type locality, Nepal. The female holotype at NHMUK is the only specimen among a collection of ~ 50 specimens, half of them originating from NE India, Assam-Meghalaya region, the second half from Japan. The volumes 1–5 of the “ Moths of Nepal ” - series do not mention it and several thousand specimens from many localities in Nepal at ZFMK definitely do not contain it. The doubt about the correctness of the type locality has arisen, but was answered by the fact that a second specimen was found at the ZSM collection, Munich, a female collected near Kathmandu, the specimen mentioned above ( Stüning 2000: 110, Moths of Nepal, vol. 6). Unfortunately, this and a male from Khasi Hills, Meghalaya could be studied by images only (Figs 2–5 ). However, in our experience they will not affect our taxonomic conclusions when investigated more closely in future.

Diagnosis.

Images of adults and genitalia of the nominate subspecies are shown here for comparison with E. languidata incrassata stat. nov. and E. yakushimensis stat. nov. The diagnoses are given under each of the two taxa.

Distribution.

Nepal (type-locality), India (Assam-Meghalaya), N. Vietnam, China ( Sichuan, Yunnan, Jiangxi, Zhejiang, Fujian, Taiwan, Guangdong, Guangxi), Korea, Japan.

Remarks.

Euryobeidia languidata is the type-species of Euryobeidia , designated by Fletcher (1979), and a member of the white or grayish-white group of species (1 st Group). It is the species with the widest area of distribution, from east Nepal in the west to north Japan in the east, with southernmost occurrence on the island of Hainan ( ssp. incrassata ). Within China, it appears to be primarily distributed along the southeastern coast. More to the north, there appears to be a gap between northern Zhejiang and South Korea, but this region is mostly low-lying plains with high population density, extensive agriculture and hardly any forests, obviously unsuitable for the existence of this species.

Figures

Figures 2–19. Adults of Euryobeidia species. 2–8. E. languidata (Walker): 2. The holotype of Abraxas languidata, female, Nepal, NHMUK; 3. Female, Nepal, upperside, ZSM; 4. Ditto, underside; 5. Male, Khasi Hills, India, ZFMK; 6. Male, N. Vietnam, CKY; 7. Male, Tokyo, Japan, CRS / NIAES; 8. Female, Niigata, Japan, CRS / NIAES; 9. E. ellipsoidea, untypical pattern, female, Sichuan, China, ZFMK; 10–13. E. languidata incrassata Xiang & Han, stat. nov., Hainan Is., China: 10. The holotype of E. incrassata Xiang & Han, male, IZCAS; 11. Non-type, male, upperside, CRICATAS; 12. Ditto, underside; 13. Non-type, female, CRICATAS; 14, 15. E. supercostata sp. nov., holotype, male, Hainan, China, IZCAS: 14. Upperside; 15. Underside; 16, 17. E. yakushimensis Inoue, stat. nov., Yakushima Is., Japan: 16. The holotype of E. languidata yakushimensis Inoue, male, NHMUK; 17. Non-type, female, CRS / NIAES; 18. E. ellipsoidea, paratype, female, Sichuan, China, ZFMK; 19. E. xuei sp. nov., paratype, male, Fansipan, Vietnam, ZFMK. Scale bar: 10 mm.

Figures 38–43. Male genitalia of Euryobeidia species. 38, 39. E. languidata (Walker): 38. Fujian, China, gen. prep. no. CRICATAS 00283; 39. Tokyo, Japan, gen. prep. no. RS 8877; 40, 41. Euryobeidia languidata incrassata Xiang & Han, stat. nov., Hainan Is., China: 40. The holotype of E. incrassata Xiang & Han, IOZ-CAS slide no. Geom- 7235; 41. Non-type, gen. prep. no. CRICATAS 00220; 42, 43. E. yakushimensis Inoue, stat. nov., Yakushima Is., Japan, gen. prep. nos. RS 7407, RS 7411. Scale bars: 1 mm.

Figures 50–55. Female genitalia of Euryobeidia species. 50. E. languidata (Walker), Japan, gen. prep. no. 2460 - DS; 51, 52. E. languidata yakushimensis Inoue, Yakushima Is., Japan, gen. prep. nos. RS 7412, RS 8879; 53. Euryobeidia languidata incrassata Xiang & Han, stat. nov., Hainan Is., China, gen. prep. no. CRICATAS 00223; 54. E. tigratoides sp. nov., paratype, Hainan Is., China, gen. prep. no. CRICATAS 00244; 55. E. tigratoides leopardiformis subsp. nov., paratype, W. Hubei, China, gen. prep. no. 2450 - DS. Scale bars: 1 mm.

Figures 56–64. Living imagos of Euryobeidia species in resting position. 56. E. languidata (Walker), Fujian, China; 57. E. languidata incrassata Xiang & Han, stat. nov., Hainan, China; 58. E. xuei sp. nov., Yunnan, China; 59. E. largeteaui (Oberthür), Guizhou, China; 60. E. quadrata Xiang & Han, Hong Kong, China; 61. E. quadrata Xiang & Han, Anhui, China; 62. E. tigratoides leopardiformis subsp. nov. or E. largeteaui, Guangdong, China; 63. E. tigratoides sp. nov., male, Hainan, China; 64. E. tigratoides sp. nov., female, Hainan, China.