Hedysarum sennoides subsp. sennoides

Adema, F., 2024, The genus Ormocarpum (Fabaceae - Papilionoideae) in Malesia and the Pacific, Blumea 69, pp. 27-35 : 33

subsp. sennoides

Shrubs, up to 4 m tall. Twigs terete, c. 5 mm diam, glabrous. Stipules (narrowly) triangular, 4.7–1.5 by 1–5 mm, both sides glabrous, caducous. Leaves with 11–13 leaflets. Petiole 4 –7 mm long, ± smooth, very shortly thinly sericeous, sometimes also with bulbous-based hairs; rachis mostly as the petiole, 30– 35 mm long; pulvinus 1–2 mm long. Leaflets: terminal one elliptic to obovate, 6.5–11 by 3.5–6 mm, 1–3.2 times as long as wide, base cuneate, apex apiculate, upper surface with very few short whitish hairs, lower surface glabrous, midrib and nerves ± sunken above, nerves 4–5 per side, 0.6–2.2 mm apart, vena- tion brownish, ± similar in colour to slightly darker than the rest of the surface; lateral leaflets mostly as the terminal one, elliptic, 7.5–9 by 3.5–3.7 mm, 2.1–2.5 times as long as wide, base obtuse; pulvinus 0.2–0.5 mm long. Bracts to the flowers orbicular, c. 0.5 by 0.4 mm, both sides glabrous. Pedicels 4–10 mm long, thinly sericeous, sometimes with bulbous-based hairs. Bracteoles broadly obovate to orbicular, 1–2.5 by 0.7–2 mm, both sides glabrous. Calyx pale green, 6.4–9 mm long, tube 6–8 mm long; teeth of upper lip triangular to semicircular, 0.5–2 by 0.5–3 mm; lateral teeth triangular, 2–3 by 1.2–3 mm, median tooth triangular, 3–5 by 2.2–4 mm; both sides glabrous. Corolla white to cream. Standard: claw 2 – 3 mm long; blade orbicular, 9–12 by 8–14 mm, apex rounded to emarginate, with 2 basal callosities, both sides glabrous. Wings: claw 3–3.2 mm long; blade obovate, 7.5–11 by 5.5–7 mm, apex rounded, outside sculptured, both sides glabrous, auricles 0.2–0.3 mm long. Keel petals: claw 3–5.5 mm long; blade ‘boat-shaped’, 8–12 by 5–7 mm, apex rounded, auricles 0.2–0.4 mm long, both sides glabrous. Stamens monadelphous, all in an open sheath or diadelphous 5 + 5, rarely 9 + 1, tube 6–10 mm long, free part 2–6 mm long, glabrous; anthers 0.7–1.1 by 0.2– 0.5 mm, glabrous. Disc 0.3–1 mm long, glabrous. Ovary 4–8.5 mm long, with bulbous-based hairs; stipe 2–3.5 mm long, glabrous; ovules 3 or 4; style 2–10 mm long, glabrous. Pods ± strap-like, c. 28 by 6 mm, articles flattened ellipsoid, 11–14 by 6–7 mm, veined, with bulbous-based hairs (see note 2). Seeds flattened ellipsoid, c. 2.8 by 1.2 by 0.9 mm; hilum c. 0.5 mm long.

Distribution — Africa; India; Sri Lanka; Malesia: Java, Philip- pines, Lesser Sunda Islands (Bali, Sumba, Sumbawa, Timor); Vanuatu.

Habitat & Ecology — Semi-dry forest , slope of mountains, along dry riverbeds, savanah.Altitude: up to 200 m. Flowering: April to August, November and December; fruiting: May, June .

Notes — 1. Willdenow (1802) cites no specimens with his descripition. In the Willdenow herbarium at B there is a speci- men collected by Klein in India annotated by Willdenow. This specimen is selected as neotype .

2. The thin hair-like part of the bulbous-based hairs may break off, leaving the bulbous base behind. The petiole, rachis, pedicel and pods may be (look) rough by these hair bases.

3. Morrison s.n. from Vanuatu with slightly smaller flowers and clearly sculptured wings has been included here.

4. Gillett (1966) recognized three subspecies: subsp. sennoides , which is widespread; subsp. hispidum (Willd.) Brenan & J.Léonard , which is found in W and C Africa and Sri Lanka and subsp. zanzibaricum Brenan & J.B.Gillett , which is found in Kenya and Tanzania.