Photuris falli Barber, 1951

Keller, Oliver & Hinson, Kevin R., 2023, Taxonomic notes on the Ototretinae, Photurinae, and Psilocladinae (Coleoptera: Lampyridae), Insecta Mundi 2023 (987), pp. 1-10 : 3-8

Photuris falli Barber, 1951

Photuris flavicollis Fall, 1927: 210 ; Lloyd 2018: 435; junior primary homonym of Photuris telephorinus flavicollis Olivier,

1886 Photuris brunnipennis var. falli Barber, 1951: 24 ; replacement name for Photuris flavicollis Fall, 1927 ; McDermott 1966:

89; 1967: 107

Note. Olivier (1886: 212) described the subspecies Photuris telephorinus flavicollis Oliver, 1886 , that was later moved into the genus Pyrogaster Motschulsky, 1853 by Olivier (1910). Later on, Fall (1927: 210) described the species Photuris flavicollis Fall, 1927 . Barber (1951: 24), acknowledging the presence of a primary homonymy, proposed falli as a new replacement name for flavicollis Fall, 1927 , which is a junior primary homonym of P. flavicollis Olivier, 1886 .

Determination of Subspecific or Infrasubspecific Rank

We evaluated all remaining taxa described as variations in the subfamilies Ototretinae , Photurinae, and Psilocladinae ( Wittmer 1944; McDermott 1966) and determined their availability based on Article 45.6 of the Code ( ICZN 1999) using Lingafelter and Nearns (2013) for elucidation of this article. In the table below, we show the taxa that have been elevated to subspecies level or were made unavailable. The table is first organized alphabetically by McDermott’s (1966) catalog, Pic’s (1924a, d) uncatalogued species, and Wittmer’s (1944) catalog, then alphabetically by subfamily and species within each subfamily in the original combination column.

OTOTRETINAE

Ototreta albicornis v. lateobscura Ototreta albicornis var. lateobscura Drilaster albicornis lateobscurus Pic, 1921: 15 Pic, 1943: 9 ( McDermott 1966: ( Pic, 1921)

118) For the corrected date of publication, see above.

Ototreta atriceps v. atricollis Pic , Ototreta atriceps var. atricollis Pic , Drilaster atriceps atricollis ( Pic, 1937a: 139 1937a: 139 ( McDermott 1966: 118) 1937)

Ototreta atriceps v. basipennis Pic , Ototreta atriceps var. basipennis Drilaster atriceps basipennis ( Pic, 1937a: 139 Pic, 1937a: 139 ( McDermott 1966: 1937)

118)

Ototreta atronotata v. pieli Pic , Ototreta atronotata var. pieli Pic , Drilaster atronotatus pieli ( Pic, 1955: 2 1955: 2 ( McDermott 1966: 118) 1955)

Ototreta corporaali v. diversipes Pic , Ototreta corporaali var. diversipes Drilaster corporaali diversipes ( Pic, 1937a: 139 Pic, 1937a: 138 [sic] (McDermott 1937)

1966: 119)

Ototreta debilis v. holzi Pic, 1914b : Ototreta debilis var. holtzi Pic , Drilaster debilis holzi (Pic, 1914)

9 1914b: 9 ( McDermott 1966: 119) For correct spelling, see above.

PHOTURINAE

Bycellonycha [sic] collaris v. bicolor- Bicellonycha collaris var. bicoloripes Bicellonycha collaris bicoloripes Pic ,

ipes Pic, 1935b: 9 Pic, 1935a [sic]: 9 (McDermott 1935 1966: 94) McDermott (1966) cites the wrong publication by Pic (1935a) in the Arkiv för Zoologi for the primary description. Bycellonycha Pic, 1935 ( Pic, 1935b: 9) is an incorrect subsequent spelling of Bicellonycha Motschulsky not in prevailing use and therefore an unavailable name.

Daiphoturis bicoloripes v. reductus Photuris ( Daiphoturis) bicoloripes Photuris ( Daiphoturis) bicoloripes

Pic, 1926: 28 var. reducta ( Pic, 1926: 28) (Mc- reducta ( Pic, 1926) Dermott 1966: 93) McDermott (1966) demoted Daiphoturis Pic, 1926 to a subgenus of Photuris Dejean.

Original combination Name used in McDermott 1966 Current status, comments

Daiphoturis bicoloripes v. suboblit- Photuris ( Daiphoturis) bicoloripes Photuris ( Daiphoturis) bicoloripes eratus Pic, 1926: 28 var. subobliterata Pic, 1926: 28 subobliterata Pic, 1926

( McDermott 1966: 93) McDermott (1966) demoted Daiphoturis Pic, 1926 to a subgenus of

Photuris Dejean.

Photuris amoena var.? nitida Gor- Bicellonycha amoena var. nitida Bicellonycha amoena nitida ( Gorham, 1884: 275 ( Gorham, 1884: 275) (McDermott ham, 1884)

1966: 94)

Photuris brunnipennis var. falli Photuris brunnipennis var. falli Photuris flavicollis Fall, 1927

Barber, 1951: 24 Barber, 1951: 24 (McDermott For discussion on the validity of 1966: 89) the name, see “Validity of Taxon Name” section above. Lloyd (2018: 435) gives Photuris flavicollis species status.

Photuris corvus v. innotata Pic , Photuris corvus var. innotata Pic , Photuris corvus innotata Pic, 1930 1930: 80 1930 a: 80 ( McDermott 1966: 89)

Photuris elegans var. guttata Ol- Pyrogaster elegans var. guttatus Pyrogaster elegans guttatus (Olivier, ivier, 1886: 215 ( Olivier, 1886: 215) (McDermott 1886)

1966: 96)

Photuris elegans var. obscuripennis Pyrogaster elegans var. obscuri- Pyrogaster elegans obscuripennis Olivier, 1886: 215 pennis ( Olivier, 1886: 215) ( Olivier, 1886)

( McDermott 1966: 96)

Photuris lucidicollis var. sordida Bicellonycha lucidicollis var. sordida Bicellonycha lucidicollis sordidus Gorham, 1884: 275 ( Gorham, 1884: 275) (McDermott ( Gorham, 1884)

1966: 95)

Photuris lunifera var. biguttata Pyrogaster lunifer var. biguttatus Pyrogaster lunifer biguttatus (Oli- Olivier, 1886: 220 ( Olivier, 1886: 220) (McDermott vier, 1886)

1966: 97) For correct spelling of the species epithet, see above.

Photuris ruficeps var. basalis Oli- Bicellonycha ruficeps var. basalis Bicellonycha ruficeps basalis ( Olivier, 1886: 242 ( Olivier, 1886: 242) (McDermott vier, 1886)

1966: 94)

Olivier (1907) synonymized Photuris ruficeps and its varieties under Photuris dimidiata (Blanchard) .

Olivier (1910) moved it to Bicellonycha .

Photuris ruficeps var. humeralis Bicellonycha ruficeps var. humeralis Bicellonycha ruficeps humeralis Olivier, 1886: 242 ( Olivier, 1886: 242) (McDermott ( Olivier, 1886)

1966: 94) Olivier (1907) synonymized Photuris ruficeps and its varieties under Photuris dimidiata (Blanchard) .

Olivier (1910) moved it to Bicellonycha .

Original combination Name used in McDermott 1966 Current status, comments

Photuris ruficeps var. rubiginosa Bicellonycha ruficeps var. rubiginosa Bicellonycha ruficeps rubiginosa

Olivier, 1886: 242 ( Olivier, 1886: 242) (McDermott ( Olivier, 1886)

1966: 94) Olivier (1907) synonymized Photuris ruficeps and its varieties under Photuris dimidiata (Blanchard) .

Olivier (1910) moved it to Bicellonycha .

Photuris telephorina var. apicalis Pyrogaster telephorinus var. apicalis Pyrogaster telephorinus apicalis Olivier, 1886: 214 ( Olivier, 1886: 214) (McDermott ( Olivier, 1886)

1966: 98)

Photuris telephorina var. atrata Pyrogaster telephorinus var. atratus Pyrogaster telephorinus atratus Olivier, 1886: 213 ( Olivier, 1886: 213) (McDermott ( Olivier, 1886)

1966: 98)

Photuris telephorina var. bicolor Pyrogaster telephorinus var. bicolor Pyrogaster telephorinus bicolor Olivier, 1886: 213 ( Olivier, 1886: 213) (McDermott ( Olivier, 1886)

1966: 98)

Photuris telephorina var. biplagiata Pyrogaster telephorinus var. Pyrogaster telephorinus biplagiatus Olivier, 1886: 213 biplagiatus ( Olivier, 1886: 213) ( Olivier, 1886)

( McDermott 1966: 98)

Photuris telephorina var. dimidiata Pyrogaster telephorinus var. Pyrogaster telephorinus dimidiatus Olivier, 1886: 213 dimidiatus ( Olivier, 1886: 213) ( Olivier, 1886)

( McDermott 1966: 98)

Photuris telephorina var. divisa Pyrogaster telephorinus var. divisus Pyrogaster telephorinus divisus Olivier, 1886: 212 ( Olivier, 1886: 212) (McDermott ( Olivier, 1886)

1966: 98)

Photuris telephorina var. flava Pyrogaster telephorinus var. flavus Pyrogaster telephorinus flavus Olivier, 1886: 213 ( Olivier, 1886: 213) (McDermott ( Olivier, 1886)

1966: 98)

Photuris telephorina var. flavicollis Pyrogaster telephorinus var. Pyrogaster telephorinus flavicollis Olivier, 1886: 212 flavicollis ( Olivier, 1886: 212) (Mc- ( Olivier, 1886)

Dermott 1966: 98)

Photuris telephorina var. intersepta Pyrogaster telephorinus var. Pyrogaster telephorinus interseptus Olivier, 1886: 213 interseptus ( Olivier, 1886: 213) ( Olivier, 1886)

( McDermott 1966: 98)

Photuris telephorina ab. mixta Pyrogaster telephorinus var. mixtus Originally described as an aberra- Olivier, 1896: 3 ( Olivier, 1896: 3) (McDermott tion, therefore automatically

1966: 98) infrasubspecific and unavailable.

Photuris telephorina var. puncticol- Pyrogaster telephorinus var. Pyrogaster telephorinus puncticollis lis Olivier, 1886: 213 puncticollis ( Olivier, 1886: 213) ( Olivier, 1886)

( McDermott 1966: 98)

Pyrogaster bilineatus v. discomacu- Pyrogaster bilineatus var. discoma- Pyrogaster bilineatus discomaculalatus Pic, 1928: 59 culatus Pic, 1928: 59 (McDermott tus Pic, 1928

1966: 96)

Pyrogaster hecticus v. atriventris Pyrogaster hecticus var. atriventris Pyrogaster hecticus atriventris Pic, Pic, 1924c: 89 Pic, 1924c: 89 ( McDermott 1966: 96) 1924

OTOTRETINAE

Gorhamia depressithorax v. testacei- Gorhamia depressithorax var. testa- Gorhamia depressithorax testaceipennis Pic, 1923: 54 ceipennis Pic, 1923: 54 (Wittmer pennis Pic, 1930

1944: 211)

Mimophaeopterus wittmeri ab. Falsophaeopterus ( Mimophae- Originally described as an aberramaculata Wittmer, 1939: 23 opterus) wittmeri ab. maculata tion, therefore automatically ( Wittmer, 1939: 23) (Wittmer infrasubspecific and unavailable 1944: 213)

Ototretadrilus flavoscutellatus ab. Ototretadrilus flavoscutellatus ab. Originally described as an aberrainnotaticollis Wittmer, 1938a: 39 innotaticollis Wittmer, 1938a: 39 tion, therefore automatically

( Wittmer 1944: 21) infrasubspecific and unavailable.

Ototretadrilus philippinus ab. Ototretadrilus philippinus ab. Originally described as an aberramaculata Wittmer, 1941: 197 maculata Wittmer, 1941: 197 (Wit- tion, therefore automatically tmer 1944: 214) infrasubspecific and unavailable.

Picodrilus drescheri var. ornaticollis Picodrilus drescheri var. ornatico- Picodrilus drescheri ornaticollis Wittmer, 1938b: 210 llis Wittmer, 1938b: 210 (Wittmer Wittmer, 1938

1944: 214)

Other nomenclatural issues