Goniofusus brasiliensis ( Grabau, 1904 )

Simone, Luiz Ricardo L., 2024, New species, misidentifications and problematic taxonomy of some Atlantic South American marine mollusks: a review, Papéis Avulsos de Zoologia 64, pp. 1-104 : 34-36

publication ID	https://doi.org/10.11606/1807-0205/2024.64.031
persistent identifier	https://treatment.plazi.org/id/4008878E-FFE8-A807-8B46-D956FAC2FD81
treatment provided by	Felipe
scientific name	Goniofusus brasiliensis ( Grabau, 1904 )
status

Treatment

Goniofusus brasiliensis ( Grabau, 1904) View in CoL ( Figs. 22-26 View Figure 22 View Figure 23 View Figure 24 View Figure 25 View Figure 26 )

Fusus brasiliensis Grabau, 1904: 65 View in CoL , 66-68 (pl. 4 figs. 1-4); Lyons & Snyder, 2019: 16, 21, 34.

Fusinus brasiliensis View in CoL : Rios, 1970: 97 (pl. 29), 1975: 104 (pl. 30, fig. 442), 1985: 108 (pl. 37, fig. 474); Dornellas & Simone, 2011: 8 (figs. 146-147); Couto et al., 2016: 311; Petuch & Berschauer, 2016: 262, 263; Lyons & Snyder, 2019: 20.

Fusinus marmoratus View in CoL : Morretes, 1949: 100; Rios, 1970: 97 (pl. 28), 2009 (fig.); Couto & Simone, 2019: 43-44 (figs. 1H, 16B, 24A, 47C) (non Philippi, 1851).

Goniofusus brasiliensis View in CoL : Vermeij & Snyder, 2018: 62-63, 67; MolluscaBase, 2023.

Types: Holotype MCZ 945 About MCZ . Paratypes: BRAZIL *, MCZ 950 About MCZ , 945 About MCZ a, 946, 947,948, 949, 950a, MZSP 18126 View Materials . Rio de Janeiro ; off Cabo Frio,35 fms, MCZ 961 About MCZ . (* inferred by the name) .

Type locality: Brazil *.

Diagnosis: Shell up to 100 mm. Spire wide, ~43° of angle; axial sculpture usually ending in penultimate whorl. Whorls shouldered, with weak shoulder in last whorl.

Differential redescription: Shell about 90 mm. Elongated, ~2.7 times longer than wide; spire~41% of total length, last whorl ~53% of total length. Spire angle ~43°. Spire up to 9 convex whorls; suture well-marked, with angle ~110°. Protoconch not preserved (eroded). Sculpture 9-10 low spiral cords, of rounded profile, interspaces equivalent to their width; additionally strong, wide axial undulations (~ 12 in penultimate whorl), being taller at middle, in which spiral cords forming blunt beaks; axial undulations becoming weak in some specimens ( Fig. 23C, D View Figure 23 ), very strong in others ( Figs. 22A, B, I, J, N, O View Figure 22 , 23A, B View Figure 23 ).Transverse section circular ( Fig. 22D, H, L, P View Figure 22 ). Last whorl with ~15 spiral cords similar to those of preceding whorls; same sculpture along siphon, as oblique cords ( Figs. 22E, I, M View Figure 22 , 23D, F View Figure 23 ). Some axial undulations in last whorl as scars of old apertures ( Figs. 22D, G View Figure 22 , 23D, F View Figure 23 ). Aperture elliptic, ~1.5 limes longer than wide; anal notch almost absent ( Figs. 22A, E, I, M View Figure 22 , 23A, C, E View Figure 23 ). Canal relatively cylindric, straight, with 30% of shell length, and ~30% of last whorl width in its base. Outer lip simply arched, weakly sinuous because of spiral sculpture; inner lip weakly concave, callus narrow, thin, not exceeding apertural limit. Umbilicus closed ( Figs. 22I, M View Figure 22 , 23A, C View Figure 23 ) to very narrowly opened ( Fig. 22A, E View Figure 22 ).

Operculum ( Fig. 23G, H View Figure 23 ): corneous, dark brown. Elliptic, with inferior terminal nucleus, on terminal beak. Inner scar occupying ~85% of inner area, with well-marked concentric, equidistant lines initiated from middle level of outer edge. Outer and interior edges thicker than remaining edges.

Head ~half of head-foot width ( Fig. 24A View Figure 24 ); cephalic tentacles (te) rather triangular, less than ⅔ of head width, distal end clearly narrower than basal region. Head flap-like ( Fig. 25A View Figure 25 ). Females with well-developed, deep cement gland ( Fig. 24B View Figure 24 : cg). Siphon stubby, very muscular ( Fig. 24C View Figure 24 : si). Gill ( Fig. 24C View Figure 24 : gi) slightly broader than osphradium (os), ~30% longer, with isosceles filament ( Fig. 24D View Figure 24 ). Osphradium symmetric ( Fig. 24D View Figure 24 : os). Proboscis very long, slender ( Fig. 25B, D View Figure 25 : pb). Rhynchostome as small pore ( Fig. 25A View Figure 25 : ry). Odontophore elongated, similar to preceding species ( Fig. 26A, B View Figure 26 ), except for bifurcation of m11 being more anterior ( Fig. 26B View Figure 26 : m11).

Radula ( Fig. 23I, J View Figure 23 ) similar to that of G. phoenix , distinctions following. Rachidian having only pair of terminal, small cusps,and broad basal reinforcement.Lateral teeth slightly less arched, having less – 11-12 – cusps.

Mid esophagus with well-developed valve of Leiblein ( Fig. 25B View Figure 25 : vl). Pair of salivary gland ducts running attached to esophageal wall short distance anterior to valve ( Fig. 25B, C, F View Figure 25 : sd). Salivary aperture in postero-lateral region of dorsal surface of buccal cavity ( Fig. 25E View Figure 25 : sd). Gland of Leiblein (gl) totally glandular, of medium size, broad anteriorly, filiform posteriorly; its duct (ld) narrow. Stomach as simple curve ( Fig. 26C View Figure 26 : st), not bulged. Penis with base wide, tapering gradually up to pointed tip ( Fig. 26D View Figure 26 ), beading small terminal papilla;penis duct (pd) totally closed (tubular). Nerve ring ( Fig. 26B View Figure 26 : nr) similar to preceding species.

Distribution: Espírito Santo to Santa Catarina coasts.

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
Family	Fasciolariidae
Genus	Goniofusus