Urochloa brizantha (Hochst. ex A.Rich.) R.D.Webster

Oulo, M., Zon, A. P. M. van der & Sosef, M. S. M., 2023, New combinations in and typification of tropical African species of Urochloa (incl. Brachiaria) (Poaceae), Blumea 68 (2), pp. 73-84 : 75

Urochloa brizantha (Hochst. ex A.Rich.) R.D.Webster

Urochloa brizantha (Hochst.ex A.Rich.) R.D. Webster (1987) 233. — Panicum brizanthum Hochst.ex A.Rich.(1850) 363. — Brachiaria brizantha (Hochst. ex A.Rich.) Stapf (1919) 531. — Lectotype (collection designated by Veldkamp 1996,duplicate designated here): Schimper 89 (lecto P [P00442084]; isolecto B?, BM [BM000923191], BR [BR0000008645142, BR000000 - 8366955],FI [FI001039],G [G00015881,G00015882,G00015884],GOET [GOET006080], HOH [HOH008887], K [K000282126, K000282127], L [L0043836, L0043837], LG [LG0000090036057, LG0000090036200], M [M0103976, M0103977], MO [MO-1742002], MPU [MPU024477, MPU-024478], P [P00442082, P00731455, P02284802, P02284803], PRE [PRE0664145-0],REG [REG000276], S [S08-12407], STU [STU000125], US [ US 00133117]), Abyssinia, plantae Adoënses, Scholoda, 3 Oct. 1837.

Panicum brizanthum Hochst. ex A.Rich. var. latifolium Oliv. (1875) 170. — Type: Speke & Grant s.n. (holo K [K000282129]), Uganda,Unjoro Plateau, Nov. 1862.

Panicum brizanthum Hochst. ex A.Rich. var. polystachyum De Wild. & T.Durand (1900) 3. — Lectotype (designated here): Thonner 78 (lecto BR [ BR0000024935609 ];isolecto BR [BR0000018400601,BR0000018400618, BR0000024935616 ]), Democratic Republic of the Congo,Yabosumba (près de Ndobo), 11 Sept. 1896.

Panicum brizanthum Hochst. ex A.Rich. var. lasiochloa Chiov. (1919) 65. — Lectotype (designated here): Bovone 75 (lecto TO), Zaire, Katanga, Plateaux des Bianos, 1800 m, Feb. 1916.

Brachiaria gangangalaensis Vanderyst (1925) 664. — Lectotype (designated by Sosef 2016): Vanderyst 5000 (lecto BR [BR0000024935593]), Democratic Republic of the Congo, Jardin agrost. Leverville, Jan. 1919.

Panicum brizanthum Hochst.ex A.Rich.var. pandanifolium Peter (1929) 31. — Type: Peter 35122 (holo B?, see notes), Tanzania, Tabora District, SE of Goweko.

Brachiaria brizantha ( Hochst. ex A.Rich.) Stapf var. angustifolia Stent & J.M.Rattray (1933) 23. — Lectotype (designated here): Eyles 1921 (lecto K [K000282057]; isolecto US [ US 00902238]), Zimbabwe, Salisbury, Nov. 1919.

Distribution — Native to most of tropical and southern Africa; introduced in most other tropical regions of the world.

Nomenclatural notes — Although Veldkamp (1996) designated the duplicate at P as the lectotype of Panicum brizanthum , it turned out that no less than 5 sheets are available, some of which appeared to have been in other, private, herbaria before. We selected one of them, likely to have been available to Richard, as the lectotype; the others are best regarded as duplicates, and hence isolectotypes.

The protologue of Panicum brizanthum var. lasiochloa mentions two syntypes, Bovone 29 and 75. At TO, a sheet carrying both specimens was located, of which the plant representing Bovone 75 has more leaves and inflorescences, and hence was chosen as the lectotype .

The holotype of Panicum brizanthum var. pandanifolium was probably lost at B during the WWII fire, and no isotypes have as yet been located, although we checked a fair number of herbaria likely to have such. Because we did not perform an exhaustive search, an isotype might still be present elsewhere, reason why we have refrained from designating a neotype.

The protologue of Brachiaria brizantha var. angustifolia mentions a large number of specimens, all to be regarded as syntypes. Not one of these has ever, as far as we could check, been assigned as the lectotype. Since the collection Eyles 1921 was the only one explicitly mentioned in both the grass treatment for Flora of tropical East Africa ( Clayton & Renvoize 1982) and Flora Zambesiaca ( Clayton 1989), and is available on-line through the JSTOR Global Plants facility (at https:// plants.jstor.org), we have chosen it as the lectotype.