Scotolathys Simon, 1885

Montana, Katherine O., Cala-Riquelme, Franklyn, Crews, Sarah C., Gorneau, Jacob A., Al-Jamal, Amin M., Alequín, Luigie D., Spagna, Joseph C., Ballarin, Francesco & Esposito, Lauren A., 2025, Tailor’s drawer no more: a reappraisal of the spider family Dictynidae O. Pickard-Cambridge, 1871 sensu lato, Zoological Journal of the Linnean Society 204 (2) : -

Genus Scotolathys Simon, 1885 s.s.

Type species: Scotolathys simplex Simon, 1885 .

Material examined: Lathys delicatula : USA, New Mexico, Bernalillo Co., Tijeras, Tunnel Canyon Trail , 7.iii.2021, Coll. M. Pleisher (1 male, CASENT9087234 ) . USA, New Mexico, Otero Co., Tularosa, 3 Rivers , 9.x.2020, Coll. M. Pleisher (1 female, CASENT9087235 ). Lathys maculina : USA, Alabama, Mobile , c. 1902, Coll. Hugo Soltau, Nathan Banks, (Holotype, 1 female, MCZ-IZ21934 ) . USA, Florida, Gainesville, Newman Lake , 16.vi.1935, Coll. W.J. Gertsch (1 male, 2 females, MNHN-AR-AR426) . USA, New Hampshire, Lake Winnepesaukee , Coll. N. Banks (1 male, 1 female, MNHN-AR-AR445). Lathys pallida (Marx, 1891) : Canada, Ontario, Point W. Bay, Lake Temagami , 15.vii.1964, Coll. W.J. Gertcsh, W. Ivie, T . B. Kurata (1 male, 2 females, MNHN-AR-AR426). Scotolathys simplex : Algeria, Oran [note: there is no further information given with this specimen] (1 female, MNHN-AR-AR426) .

Diagnosis: Scotolathys s.s. can be distinguished from other Lathyidae fam. n. by lacking AME ( Fig. 5D–F; Marusik et al. 2009: figs 1–3, 9, 28) (AME present in Lathys s.s., Analtella stat. reinst., and Andronova gen. n.); male palp without patellar apophysis ( Marusik et al. 2009: figs 33–37) (present in Lathys , Bannaella , and Denticulathys gen. n.); cymbium with a prolateral proximal process ( Marusik et al. 2009: figs 41, 42, 48) (absent in Afrolathys gen. n., Andronova gen. n., Bannaella , Denticulathys gen. n., Analtella stat. reinst., and Lathys s.s.); embolus usually coiled 1080° or less, usually with a distal retrolateral origin ( Marusik et al. 2009: Figs 39, 40, 42, 49); the conductor tip is thin, elongated, usually coiled 360°, and with a paraterminal process ( Marusik et al. 2009: figs 39, 48, 50). Females ( Marusik et al. 2009: figs 58–61) can be distinguished by the reduced SS/AG so that it is inconspicuous; copulatory duct elongated and coiled 720°or more around itself and bent 360° around the primary spermatheca; fertilization duct located mesal distally.

List of included species: Scotolathys delicatula Gertsch and Mulaik, 1936 stat. reinst., S. immaculata Chamberlin and Ivie, 1944 stat. reinst., S. maculina (Gertsch, 1946) stat. reinst., S. pallida (Marx, 1891) stat. reinst., and S. simplex (Simon, 1885) .

Genus Tolokonniella Cala-Riquelme, Crews, Esposito gen. n.

urn:lsid:zoobank.org:act: CB 5C2ACD-C071-4D40-B977-

C8F68D8EAE7D.

Type species: Lethia stigmatisata Menge, 1869 , currently Lathys stigmatisata (Menge, 1869) .

Material examined: Lathys stigmatisata (Menge, 1869) : France, Paris , 18.vi.1913, Coll. J.E. Simon (1 male, 1 female, NHMUK1635150 ) .

Etymology: The generic epithet is in honour of the Russian musician, conceptual artist, and feminist Nadezhda Andreyevna Tolokonnikova; gender feminine.

Diagnosis: The male of Tolokonniella gen. n. resembles Lathys s.s. by having the AME narrower than the ALE ( Özkütük et al. 2016: figs 2, 4); the male palp with a patellar dorsal distal apophysis ( Marusik et al. 2009: figs 29, 44, 52, Marusik et al. 2015: fig. 50, Özkütük et al. 2016: figs 8–15); palpal tibia with a ventral lateral apophysis in addition to the RTA and a hook-shaped tibial process; however, this genus can be distinguished from Lathys s.s. by having the embolus coiled more than 1080°, and usually with a proximal prolateral origin ( Marusik et al. 2015: fig. 51); the conductor tip elongated and coiled three or more times ( Marusik et al. 2006b: fig. 9). Females ( Marusik et al. 2009: figs 31, 32, Marusik et al. 2015: figs 48, 49, Özkütük et al. 2016: figs 17–21; 37–40) can be distinguished by the reduced SS/AG, the copulatory duct elongated and coiled 720° around itself, and 180° around the primary spermatheca; the fertilization duct located mesal distally.

List of included species: Tolokonniella ankaraensis ( Özkütük et al., 2016) comb. n., T. mallorcensis (Lissner, 2018) comb. n., T. maura (Simon, 1911) comb. n., T. stigmatisata (Menge, 1869) comb. n., and T. truncata (Danilov, 1994) comb. n.