Analtella Denis, 1947

Montana, Katherine O., Cala-Riquelme, Franklyn, Crews, Sarah C., Gorneau, Jacob A., Al-Jamal, Amin M., Alequín, Luigie D., Spagna, Joseph C., Ballarin, Francesco & Esposito, Lauren A., 2025, Tailor’s drawer no more: a reappraisal of the spider family Dictynidae O. Pickard-Cambridge, 1871 sensu lato, Zoological Journal of the Linnean Society 204 (2) : -

Genus Analtella Denis, 1947 , stat. reinst.

Type species: Analtella brevitarsis Denis, 1947 , currently Lathys narbonensis (Simon, 1876) .

Material examined: Lathys affinis (Blackwall, 1862) : Portugal, Madeira, Porto Santo Island , 32°44'48.8754"N 16°42'11.052"W, iv.2017, Coll. J. Malumbres-Olarte et al. (2 females, 4 imm, CASENT9081474 ) . Portugal, Madeira, Porto Santo, Pico do Castelo , 17.iv.2017, Coll. L. Crespo, I. Silva (1 female CASENT9081478 ) . Portugal, Madeira, Porto Santo, Ilheu de Ferro , 18.iv.2017, Coll. L. Crespo, I. Silva (1 female, CASENT9081479 ) . Portugal, Madeira, Deserta Grande, Planalto Sul , 11.iv.2017, Coll. L. Crespo, I. Silva (1 female, CASENT9081480 ) . Portugal, Madeira, Deserta Grande, Planalto Sul , 11.iv.2017, Coll. L. Crespo, I. Silva (1 female, CASENT9081481 ) . Portugal, Madeira, Porto Santo, Pico do Castelo , 17.iv.2017, Coll. L. Crespo, I. Silva (1 female, CASENT9081482 ) . Portugal, Madeira, Porto Santo, Rocha De Nossa Senhora , 21.iv.2017, Coll. L. Crespo, I. Silva (1 female, CASENT9081483 ) . Portugal, Madeira, Porto Santo, Pico Branco , 23.iv.2017, Coll. L. Crespo, I. Silva (1 male, CASENT9087229 ) . Lathys dentichelis : Portugal, Azores, Terceira Island, Lagoinha , Coll. Project BALA (1 male, 2 females, CASENT9081472 ; 1 female, CASENT9081473 ) . Equatorial Guinea, Bioko, Moca, 3°21′46′′N, 8°39′52′′E, Elev. 1400 m, 4–9.x.1998, Coll. D.K. Dabney, D. Ubick (1 female, CASENT9118700 ) .

Diagnosis: The male of Analtella stat. reinst. ( Figs 13C–F, 38AF) resembles Andronova gen. n. by having three apophyses that block the distal end of the conductor ( Bosmans and Gavalas 2023: fig. 1E, H); however, it can be distinguished from Andronova gen. n. and other Lathyidae fam. n. by the chelicerae with an ectal, longitudinal row of macrosetae ( Fig. 38A; Bosmans and Gavalas 2023: fig. 1B, C); the reduced AME ( Fig. 38A; Bosmans and Gavalas 2023: fig. 1B) (AME absent in Afrolathys gen. n., Scotolathys s.s.) embolus usually coiled less than 1080°, and usually with a retrolateral distal origin; palpal tibia with lamellate lateral apophysis with deep or shallow longitudinal furrow in addition to the RTA, and hook-shaped tibial process (lamellate lateral apophysis absent in Lathys s.s., Scotolathys s.s., Bannaella , Afrolathys gen. n.); the conductor tip elongated, not compressed, and coiled two or more times ( Fig. 38F). Females ( Fig. 13C–F; Bosmans and Gavalas 2023: fig. 1I, J) can be distinguished by the SS/AG, usually shorter than the primary spermathecae; copulatory duct elongated and coiled more than 720° around itself and the SS/AG, and bent 90° or less around the primary spermatheca; the fertilization duct is located mesal distally.

Remarks: Analtella was described by Denis (1947: 145) based on females of Analtella jubata Denis, 1947 ( 1947:146) and Analtella brevitarsis Denis, 1947 ( 1947: 148) in a comparison with species of Altella . Denis (1947) did not designate a type species, so Lehtinen (1967: 243) designated A. brevitarsis as the type of Analtella . In the same paper, Lehtinen (1967: 243) considered A. brevitarsis a junior synonym of Lathys narbonensis Simon, 1914 and placed Analtella and Scotolathys within the narbonensis group, while A. jubata (= Lathys jubata ) became part of the puta group with L. stigmatisata . Here, we reinstate the status of Analtella to include L. narbonensis Simon, 1914 based on the morphology of

62 • Montana et al.

the male palp and the female genitalia. Additionally, we consider L. jubata a junior synonym of L. stigmatisata (= Tolokonniella stigmatisata comb. n.) based on the morphology of the epigynum with reduced SS/AG, the copulatory duct elongated and coiled 720° around itself, and 180° around the primary spermathecae.

List of included species: Analtella affinis (Blackwall, 1862) comb. n., A. dentichelis (Simon, 1883) comb. n., A. narbonensis (Simon,

1876) comb. n., A. pygmaea ( Wunderlich, 2011) comb. n., and A. teideensis (Wunderlich, 1992) comb. n.