Tisamenus lachesis ( Rehn & Rehn, 1939 )

Hennemann, Frank H., 2025, A taxonomic review of Philippine Obrimini stick insects: The genus Tisamenus Stål, 1875 (Insecta: Phasmatodea: Heteropterygidae: Obriminae), Faunitaxys 13 (24), pp. 1-85 : 50-55

Tisamenus lachesis ( Rehn & Rehn, 1939)

( Fig. 27 -30, 46O-P)

Hoploclonia lachesis Rehn & Rehn, 1939: 471 .

HT, ♂: Polillo, Taylor ; Hoploclonia lachesis Rehn & Rehn , Type; Type No. 53314 U.S. N.M. [USNM].

Tisamenus polillo, Zompro, 2004: 206 .

- Otte & Brock, 2005: 335.

- Brock & Büscher, 2022: 521.

- Hennemann, 2023b: 128.

Tisamenusserratorius, Dräger, 2012: 12, figs. 16-17. (Misidentification)

- Krijns, 2011: 7. (Misidentification – culture report)

Tisamenus sp. „ Quezon National Park“, Dräger, 2012: 12, figs. 18-22.

Tisamenus sp. „Cunayan“, Dräger, 2012: 13.

Material examined

1 ♀: Coll. R.I.Sc.N.B., Philippines, E Luzon, Quirino, Sierra Madre , V.2014, local collector, I. Lumawig, [ RBINS] ;

1 ♀: Coll. R.I.Sc.N.B., Philippines, E Luzon, Quirino, vi.2014, 16°17’N 122°35’E, Leg. I. Lumawig, gift from B. Kneubühler, I.G.: 32.613 [ RBINS] ;

2 ♂: Coll. R.I.Sc.N.B., Philippines, E Luzon, Quirino, Sierra Madre , iv.2014, Purchased from I. Lumawig, Gift from B. Kneubühler, I.G.: 32.613 [ RBINS] ;

1 ♀: Coll.R.I.Sc.N.B., Philippines,Luzon, Aurora Prov., Cunayan Falls [ RBINS] ;

20 ♀, 13 ♂, 1 ♀ (penultimate instar), 20 eggs: Ex Zucht: F. Hennemann 2011-2013, Herkunft: Philippinen, Luzon Id., Prov. Quezon, Reál, 2009 [ FH, No’s 0727-1 to 34, E1] ;

4 ♀, 1 ♂, 1 ♀ (penultimate instar): Ex Zucht: F. Hennemann 2014/15, Herkunft: Philippinen, Luzon, Prov., Aurora, Cunayan Falls , 2009 (PSG 359) [ FH, No’s 0727-35 to 40] ;

5 ♀, 6 ♂, 1 ♀ (immature), 3 eggs: Philippinen, Ost Luzon Island, Provinz Aurora, Sierra Madre , Dingalan , 230 m, local collector X.2012 [ FH, No’s 0727-41 to 51, E2] ;

1 ♀, 1 ♀ (penultimate instar): Philippinen, S Luzon Id., Bicol Region, Provinz Sorsogon, Mount Bulusan , local collector IX.2012 [ FH, No’s 0727-52 & 53] ;

1 ♂: Philippinen, O Luzon Id., Prov. Isabela, Sierra Madre, Cagayan Valley,Palanan,local collector II.2011 [ FH, No.0727-55] ;

1 ♀, 1 ♂: Philippinen,S-LuzonId., ProvinzQuezon , Sierramadre, Real, local collector II.2012 [ FH, No’s 0727-58 & 59] ;

1 ♀, 2 ♂: Philippinen, S-Luzon Id., Provinz Quezon, Calabarzon , General Nakar Munip., collector XII.2012 [ FH, No. 0727-60 to 62] ;

1 ♂: Philippinen, S-Luzon, Provinz Bulacan, Santa Maria Munip., local collector V.2012 [ FH, No. 0727-57] ;

1 ♂, 3 ♀ (immature): Philippinen, OstLuzonIsland, Provinz Aurora, Sierra Madre, Dingalan , 230 m, local collector III.2012 [ FH, No’s 0727-63 to 66] ; 4 ♀, 5 ♂ 2 ♀ (immature): Philippinen, Provinz Laguna, Santa Rosa Municipality , local collector VI-VII.2012 [ FH, No’s 0727-67 to 77] ;

2 ♀ (penultimate instar): Philippinen, Provinz Nueva Vizcaya, Santa Fe Munip., Canabuan , local collector XI.2012 [ FH, No’s 0727-78 to 79] ;

1 ♀: Philippinen, Eastern Visayas, Provinz Northern Samar, Samar Island, Lope de Vega , local collector III.2012 [ FH, No’s 0727-56] .

Differentiation. – Femalesof thiswidely distributedLuzonian species ( Fig. 27) are morphologically closest to those of T. polillo ( Rehn & Rehn, 1939) , which seems to be sympatric at several of the known localities including Polillo Island, the type-locality of both species, as well as T. serratorius Stål, 1875 . These ♀ can however easily be separated fromthose of polillo by on average larger size, always having five distinct spines along the lateral margins of the mesopleurae (only three in polillo , of which only the mesopleural is frequently spinose with the other spines often only represented as tubercles), presence of 1-2 distinct metapleural laterals (none in polillo ), presence of more or less distinct paired inter-posterior meso- and metanotals ( Fig. 27G, J and 28 G-H), which however may occasionally only be represented as conicaltubercles(wholly wantingin polillo ),strongly raised and spinose anterior angle of the carinae of the triangular mesonotal area ( Fig.28 FH; just weakly protruded in polillo ) and presence of second paired posterior spinesonabdominaltergumV.From serratorius these ♀ differ by the much larger size (body length> 57.0 mm), relatively longer mesonotum, smaller triangular mesonotal area, which is no more than half as long as the mesonotum ( Fig. 28 F-H), having only five strong mesopleural spines (six distinct spines in serratorius ), notably more developed and spinose cephalic armature and body spination, and longer subgenital plate, that notably projects beyond the tip of the epiproct ( Fig. 28 A-E). Males ( Fig. 29 A-G) show the closest morphological affinity to those of polillo , but can be distinguished by the stockier overall shape and limbs, always having five strong mesopleural spines (at best four spines in polillo of which only the mesopleural is frequently spinose), presence of distinct metapleural laterals, presence of paired inter-posterior meso- and metanotals ( Fig. 29 N-O), strongly protruded and more or less spinose anterior angle of the carinae of the triangular mesonotal area ( Fig. 29 N-O; just weakly raised in polillo ) and more acute medio-longitudinal carina of the mesosternum ( Fig.29P).These ♂ also strongly resemble those of serratorius but are larger (body length> 41.0 mm) and considerably slenderer and can readily be separated by having only five instead of six mesopleural spines and having the triangular mesonotal area extending not further than to the middle of the segment. From the essentially similar but smaller and much stockier T. clotho ( Rehn & Rehn, 1939) both sexes of lachesis are readily distinguishable by the relatively longer body segments, less widened meso- and metapleurae and shorter pleural spines, as well as the more elongate mesonotal triangular area, which is notably longer than wide (about as long as wide across the anterolateral angles and roughly forming an isosceles triangle in clotho ). The eggs ( Fig. 46 O-P) differ from all other known eggs of congenerics by the more elongate shape being as much as 1.7xlonger than wide, very dense and rather uniform meshwork of densely setose ridges and short posterior lobes of the micropylar plate, which do not extend to the middle axis of the capsule. The shortposterior extensions are shared with eggs of T.hebardi ( Rehn & Rehn, 1939) but in that species the extensions are even shorter and only form two rounded lobes.

Description

♀ ( Fig. 27)

Form and colouration. – Size variable but large for the genus (body length 57.0-70.0 mm); general form quite slender and elongate; the elements of armature strongly developed and mostly spinose; legs elongate and slender with weakly developed armature. Body surface unevenly granular. General colour mostly ranging from fairly plain drab over various tones of brown and fuscous to almost black, but also clay coloured to ochraceous specimens occur A. Dorsal view (captive reared from Real, Sierra Madre, Quezon Province, Luzon ) [ FH 0727-8 ]. B. Dorsolateral view (captive reared from Real , Sierra Madre , Quezon Province, Luzon ) [ FH 0727-8 ]. C. Lateral view (captive reared from Real , Sierra Madre , Quezon Province, Luzon ) [ FH 0727-8 ]. D. Ventral view (captive reared from Real , Sierra Madre , Quezon Province, Luzon ) [ FH 0727-8 ]. E. Dorsal view (captive reared from Cunayan , Aurora Province, Luzon ) [ FH 0727-36 ]. F. Dorsolateral view (captive reared from Cunayan , Aurora Province, Luzon ) [ FH 0727-36 ]. G. Lateral view (captive reared from Cunayan , Aurora Province, Luzon ) [ FH 0727-36 ]. H. Dorsal view (from Dingalan , Aurora Province, E-Luzon) [ FH 0727-54 ]. I. Dorsolateral view (from Dingalan , Aurora Province, ELuzon) [ FH 0727-54 ]. J. Lateral view (from Dingalan , Aurora Province, E-Luzon) [ FH 0727-54 ]. K. Closeup of head, pro- and mesonotum of light brown live captive reared ♀ from Real, Quezon Province, Luzon, dorsolateral view. L. Closeup of head, pro- and mesonotum of live dark brown captive reared ♀ from Real, Quezon Province, Luzon, dorsal view .

that have distinct black V-shaped posterior markings on the meso- and metanotum and black spots in front of the second paired posteriors of abdominal terga II-IV ( Fig. 27A, E); often the area along medio-longitudinal keel of meso- and metanotum dark brown in the lighter specimens ( Fig. 27K). Meso- and metasternum lighter in colour and buff to almost clay coloured ( Fig. 27D). Femora mostly mid to dark brown and almost black, usually darker than body. Largest spines of head and body with apical half dark orange.Antennae with terminal five joints ochre to dark orange.

Head. – Basically, as in ♂; somewhat longer than wide, the eyes much smaller, sub-globose and their diameter corresponding to a little less than 0.4x the length of gena. Armature essentially as in ♂, but coronals less than half as large as the spinose supra-orbitals; the occipitals smaller than coronals, conically tubercular ( Fig. 28 F-H). Antennae almost reaching to tip of front legs and consisting of 26 joints; scapus and pedicellus like in ♂, the terminal joint almost as long as two preceding ones combined.

Thorax. – Pronotum like in ♂; the strong anteriors however basically trifid and ranging from tubercular to strongly spinose with the anterior spike anteriorly projecting over anterior margin of pronotum ( Fig. 28 F-H), but occasionally with an additional spike posteriorly ( Fig. 28H) or with single, much smaller intercalated points. Like in ♂, the transverse median sulcus notably shifted towards the posterior. Mesothorax narrowed anteriorly and very slightly widening towards posterior;2.6x longer than prothorax and 1.4x longer than prothorax and posterior portion 1.3x wider than anterior margin. Mesonotum elongate, slightly convergent towards the posterior and with a small narrowing one-quarter before posterior margin and 2x longer thanwidth at anterior margin; triangular area shorter than in ♂, scarcely attaining middle of segment and roughly an isosceles triangle, the convergent margins granular and becoming slightly tubercular towards the tubercular to spinose anterolateral angle ( Fig. 28 F-H). Posterior half of mesonotum with a distinct, granular medio-longitudinal keel that is weakly indicated in the triangular area; inter-posteriors variable in size and ranging from low, sub-obsolete tubercles to distinct spines. Mesopleurae moderately expanding towards posterior and like in ♂ with three strong, slender andlong spinose laterals, the anterior one of which is notably smaller than the three following ones; mesopleurallarge, spinose, notably larger than laterals and occasionally witha few small tuberclesaround the base. Metanotum weakly sub-trapeziform; the medio-longitudinal keel and inter-posteriors like on mesonotum. Metapleurae like in ♂ with two spinose laterals and a very strong, spinose supra-coxal at the moderately expanded angle; metapleural small, tubercular. Mesosternum distinctly tri-carinate and destitute of mesosternals ( Fig. 28I). Metasternum only with a weakly indicated medio-longitudinal carina ( Fig. 27D).

Abdomen. – Median segment distinctly trapezoidal with anterior margin rounded; the medio-longitudinal carina shallow and marked by clustered granules; occasionally with a transverse row of four small nodes at posterior margin. Segments II-VII scarcely sub-equal in length; II-III weakly increasing and V-X almost gradually and notably decreasing in width; II-VII transverse with V about 2x wider than long. All terga with two fine and closely spaced, parallel medio-longitudinal carinae which are just faintly indicated on II-IV but rather definite on following; II-V with a more or less prominent second-paired posterior spines and a variably sized posterior mesal, that varies from tubercular to distinctly spinose but is always smaller than the second-paired posteriors. Sterna II-VII with a fine medio-longitudinal carina; praeopercular organ formed by a two shallow, converging bulges that posteriorly terminate in a small swelling. Tergum VIII trapezoidal in dorsal view, IX obtusely tectate with an obtuse, triangular posteromedian protrusion formed by the two medio-longitudinal carinae ( Fig. 28 A-B). Anal segment strongly declining with the lateral margins obliquely convergent in posterior half and with a fairly well developed medio-longitudinal carina; close two anterior margin with a pair of nodes and posterior margin broadly rounded ( Fig. 28 A-D). Epiproct weakly tectate longitudinally with an indication of a medio-longitudinal carina, the lateral margins weakly convergent and the posterior margin roundly angular ( Fig. 28 C-D). Subgenital plate fairly elongate, notably projecting beyond epiproct, navicular, distinctly carinate in posterior half with the apex narrowly triangular ( Fig. 28E).

Legs. – All slender and elongate with the teeth rather weakly developed but all acutely triangular; the two apical ventral teeth of meso- and metafemora rather spinose and those of the metafemora the largest of all teeth. Profemora roughly equal in length to mesothorax, mesofemora notably shorter; metafemora reaching to posterior margin of abdominal segment V and metatibiae almost reaching to tip of epiproct. Pro- and mesotibiae wholly unarmed, metatibiae only with 5-7 small denticles or indications of teeth on twoventral carinae.Basitarsirather elongate, almostas longas following three tarsomeres taken together.

Measurements [mm]. – Body 57.0-70.0, pronotum 4.5-5.0, mesonotum 10.3-13.0, metanotum 4.7-5.8, median segment 2.4-2.8, profemora 10.3-12.5, mesofemora 9.2-11.5, metafemora 12.0-14.8, protibiae 10.2-12.0, mesotibiae 9.1-11.2, metatibiae 12.3-15.4, antennae 19.5-21.0.

Variability. – Both sexes of this species show considerable intraspecific variability in size, armature of the head and body and also some variability is seen in the colouration. While the morphological traits show a certain degree of affiliation to individual populations, the chromatic variation appears to be not much correlative to the distribution. It seems only noteworthy that light clay coloured to ochraceous ♀ with distinct black V-shaped posterior markings on the meso- and metanotum and black spots in front of the second paired posteriors of abdominal terga II-IV are predominantly found in the population at Cunayan Falls, Province Aurora ( Fig. 27 A-B, E-F). These are also found among specimens from other localities but relatively much lesser in number. While ♂ are more constant in colour and mostly varying from dark buff over sepia and umber to dark brown, often with a slight orangey wash, ♀ show more variability in colour. Aside from the colour morph described above, the majority of specimens is rather plain drab to fuscous ( Fig. 27 H-J) or even almost black. In the lighter coloured specimens, the femora are contrastive dark brown and usually these are darker than the body.

All elements of cephalic,thoracic and abdominal armature are subject to variability and vary in size. The elements with the most noteworthy variability are the pronotal anteriors, antero-laterals of the triangular mesonotal area, inter-posteriors of the meso- and metanotum and medial of abdominal terga II-V. The latter are sub-obsolete or merely represented as small tubercles in most of the ♀ examined but developed to short but distinct spines in the examples from Cunayan Falls and some of the specimens from Dingalan , Province Aurora. All ♂ from Palanan , Province Isabela, Cunayan Falls and Dingalan possess a prominent pair of spinose inter-posterior meso- and metanotals ( Fig. 27 F-G and I-J, 29F-G). These are comparatively smaller and rather tubercular in the examples from Santa Rosa , Province Laguna, whereas they are sub-obsolete and merely seen as small tubercles in specimens from other localities (e. g. Fig. 27C, 29C). The population from Dingalan is interesting in that three of the five ♀ at hand have these mesonotal and metanotal inter-posteriors well developed and sub-spinose, whereas they are sub-obsolete to almost complete missing in the other two specimens. The anterolateral mesonotals at the anterior angles of the triangular area vary from obtusely tubercular (e. g. Fig. 28F) to long and acutely spinose ( Fig.28 G-H, 29O), generally being mostdeveloped in the populations at Dingalan and Santa Rosa . The anterior pronotals are prominent and basically trifid in all ♀ at hand, but vary considerably in size and occasionally have an additional but smaller posterior spike ( Fig. 28 GH). The length of the individual tubercles or spines varies from almost uniform to distinctly unequal. In ♂ the pronotal anteriors are always very long and spinose but either bi- or trifid. The ♂ from Santa Maria , Province Bulacan ( FH, No. 0727-57) is remarkable for having the mesopleural laterals much smaller than all other examined specimens. Since only this unique specimen is available from this particular locality, it is not possible to decide whether this trait is typical for this population or merely individual .

A remarkable range is also seen in the size of this species, that varies between different populations. The holotype ♂ from the island of Polillo is larger than all other examined specimens with a body length of 51.0 mm. For summarizing the size ranges, body lengths are given here for some of the populations and are as follows: Province Quezon (Real) ♀ 59.0-70.0 mm, ♂ 42.5-48.0 mm; Province Aurora (Cunayan) ♀ 57.0-60.0 mm, ♂ 44.5 mm; Province Aurora (Dingalan) ♀ 61.5-67.0 mm, ♂ 41.5-43.0 mm; Samar ( Lope de Vega ) ♀ 60.0 mm. A full set of the size range of ♂ is given below .

A. Terminalia in lateral view (captive reared from Real,Sierra Madre, Quezon Province,Luzon) [FH 0727-8]. B. Terminalia in lateral view (from Dingalan, Aurora Province,E-Luzon) [FH 0727-54]. C. Terminalia in dorsal view (captive reared Cunayan, Aurora Province, Luzon) [FH 0727-36]. D. Terminalia in dorsal view (captive reared from Real, Sierra Madre, Quezon Province,Luzon) [FH 0727-8]. E. Terminalia in ventral view (captive reared from Real, Sierra Madre, Quezon Province, Luzon) [FH 0727-8]. F. Closeup of head,pro- and mesonotum (captive reared from Real, Sierra Madre, Quezon Province, Luzon) [FH 0727-8]. G. Closeup of head, pro- and mesonotum (from Dingalan, Aurora Province, E-Luzon) [FH 0727-54]. H. Closeup of head, pro- and mesonotum (captive reared from Cunayan, Aurora Province, Luzon) [FH 0727-36]. I. Closeup of head, pro- and mesosternum (captive reared from Real, Sierra Madre, Quezon Province, Luzon) [FH 0727-8].

Measurements [mm]. – Body 41.5-51.0, pronotum 3.0-3.5, mesonotum 8.5-10.0, metanotum 3.8-4.2, median segment 2.2-2.5, profemora 8.7-9.8, mesofemora 8.0-8.5, metafemora 9.4-10.6, protibiae 9.1-10.0, mesotibiae 8.0-8.7, metatibiae 10.3-10.9, antennae 16.5-18.0.

Egg ( Fig. 46 O-P)

Large and rather elongate for the genus; capsule ovoid, slightly oval in cross-section with the lateral surfaces almost parallel-sided in median portion, higher than wide with dorsal surface not noticeably bulgier than ventral surface, the polar-area weakly indented; capsule 1.6x longer than wide. Surface all over covered by fairly long fringy to hairy structures,that are arranged in an irregular and moderately dense meshwork; the fringes only wanting on a moderately broad rim just below anterior margin; that margin also covered by these excrescences. Micropylar plate fairly large and almost 0.8x the length of capsule; Y-shaped with the median portion roughly parallel-sided and almost approaching anterior margin of capsule, the two posterolateral extensions slender and variable in length, either not reaching axis of egg capsule or in the examples from Dingalan (coll. FH 0727-E2) slightly surpassing the axis if seen laterally. Posterior portion 90° V-shaped and with a small and shallow bowl-shaped micropylar cup in centre. Outer margin of plate marked by a rim of fringy excrescences, the inner portion sculptured like capsule and the rim in between outer margin and the slightly raised interior portion smooth and somewhat lowered. Operculum roundly conical, with an outer collar of fringy excrescences and in ventral portion with a broad rim of somewhat longer fringy to hairy structures. Colour plain brown with all the fringes of the capsule, micropylar plate and operculum dark ochre to buff. Measurements [mm]: Length incl. operculum 4.7, length 4.4, width 2.8, height 3.0, length of micropylar plate 3.5.

A. Dorsal view (captive reared from Real, Sierra Madre , Quezon Province, Luzon ) [ FH 0727-18 ]. B. Dorsolateral view (captive reared from Real , Sierra Madre , Quezon Province, Luzon ) [ FH 0727-18 ]. C. Lateral view (captive reared from Real , Sierra Madre , Quezon Province, Luzon ) [ FH 0727-18 ]. D. Ventral view (captive reared from Real , Sierra Madre , Quezon Province, Luzon ) [ FH 0727-18 ]. E. Dorsal view (from Palanan , Cagayan Valley , Isabela Province, Luzon ) [ FH 0727-55 ]. F. Dorsolateral view (from Palanan , Cagayan Valley , Isabela Province, Luzon ) [ FH 0727-55 ]. G. Lateral view (from Palanan , Cagayan Valley , Isabela Province, Luzon ) [ FH 0727-55 ]. H. Terminalia in lateral view (captive reared from Real , Sierra Madre , Quezon Province, Luzon ) [ FH]. I. Terminalia in lateral view (from Dingalan, Aurora Province, E-Luzon) [ FH]. J. Terminalia in dorsal view (captive reared from Real, Sierra Madre, Quezon Province, Luzon) [ FH]. K. Terminalia in dorsal view (Dingalan, Aurora Province, E-Luzon) [ FH]. L. Terminalia in ventral view (captive reared from Real, Sierra Madre, Quezon Province, Luzon) [ FH]. M. Anteroventral view of right hind leg (from Palanan, Cagayan Valley, Isabela Province, Luzon) [ FH 0727-55 ]. N. Closeup of head, pro- and mesonotum (captive reared from Real, Sierra Madre, Quezon Province, Luzon) [ FH 0727-18 ]. O. Closeup of head, pro- and mesonotum (captive reared from Cunayan, Aurora Province, Luzon) [ FH 0727-18 ]. P. Closeup of head, pro- and mesosternum (captive reared from Real, Sierra Madre, Quezon Province, Luzon) [ FH 0727-18 ] .

Remarks. – This species was originally described from a unique ♂ from the island of Polillo in the collection of ANSP and since a detailed description of the ♂ was presented by Rehn & Rehn (1939: 471), intraspecific variability is summarised above and the elements of armature essentially agree between the sexes,only a description of the previously unknown ♀ is presented herein. The eggs are formally described for the first time. At several known localities this species is sympatric with T.polillo ( Rehn & Rehn, 1939) .

Several stocks of T. lachesis from different localities are or have been cultured in Europe. A first stock was collected in the village Real and close to Real in the Sierra Madre mountains, Quezon province, South Luzon in 2009 by Joachim Bresseel (RBINS) and Thierry Heitzmann ( Philippines) and was the first stock of a Tisamenus -species to be introduced to Europe. Further specimens were found with additional stockintroducedto Europe by Joachim Bresseel (RBINS), Rob Krijns (Maastricht, Netherlands) and Tim Bollens (Herselt, Belgium) in 2010 ( Dräger,2012:12). This stock has been referred to as either Tisamenus sp. ‘Sierra Madre’ or Tisamenus sp. ‘Real’ was misidentified as T. serratorius and subsequently given the Phasmid Study Group culture No. 314. Already in November 2008 however, Thierry Heitzmann ( Philippines) collected a ♀ at Atimona, Quezon National Park, which is about 70 km away from Real ( Dräger, 2012: 12) and reared a stock in captivity.Stock that is offspring from this ♀ and exported to Europe has been distributed as Tisamenus sp. ‘Quezon’ or T. serratorius ‘QNP’ and seems to be still maintained by some breeders although. It has however, not been given a PSG culture number (personal communication with Holger Dräger in January 2025). A third stock was collected at the Cunayan Falls, San Luis, Aurora Province. This was distributed as Tisamenus sp. ‘Cunayan’and given culture No. 359 by the Phasmid Study Group. It seems, that only the first two stocks from Sierra Madre are still maintained in culture, while the Cunayan stock was lost after a few generations. This species has proven very easy and it has become very popular among European breeders because of the pretty colour variations.

Distribution. – Polillo [USNM – type locality]). Luzon: Province Quirino (Sierra Madre [RBINS]); Province Aurora (San Luis, Cunayan Falls [RBINS, FH]; Sierra Madre, Dingalan 230 m [FH]); Province Isabela (Sierra Madre, Cagayan Valley, Palanan [FH]); Province Quezon (Sierra Madre,Real[FH; photographic records byAlbert Kang: https://inaturalist.ca/observations/57611953 and Gernot Kunz: https:// inaturalist.ca/observations/24796645]; Sierra Madre, Real,Marikina-Infanta highway [photographic record:https://inaturalist.ca/observations/53592446]; Sierra Madre, Atimona [stock collected by T. Heitzmann in 2008]; Province Nueva Vizcaya (Santa Fe, Canabuan [FH]); Province General Nakar [FH]); Province Sorsogon (Mount Bulusan [FH]); Province Bulacan (Santa Maria [FH]); Province Laguna (Santa Rosa [FH]; Siniloan-Famy [photo by Albert Kang: https://www.jungledragon.com/ image/47185/stick_insect_phasmid_-_tisamenus _serratorius .html]). Samar: Province North Samar (Lope de Vega [FH]).