Galerita rufofemorata, Fedorenko, 2025

Fedorenko, D. N., 2025, A review of Galerita (Coleoptera: Carabidae: Dryptini) of Vietnam, with description of ten new species, Russian Entomological Journal 34 (1), pp. 37-63 : 40-41

KEY TO THE SPECIES OF GALERITA OF VIETNAM:

1(6) Body macropterous but rare exceptions (see couplet 5). Elytra evenly convex in basal third, with sides only slightly dilated apicad. Femora except black apices and often also head and pronotum pale. Pronotum 1/2–3/5 wider at base than at apex.

2(3) Head and pronotum less shiny due to a slightly coarser microsculpture. Punctation shallower, very sparse at middle of basal 1/2 head, thus leaving center of vertex nearly smooth. Apex of median lobe of aedeagus round in lateral view, without preapical ventral tooth ( Fig. 22 ). — India and Nepal to Myanmar............ [ G. orientalis Schmidt-Göbel, 1846 ]

3(2) Head and pronotum glossy, with punctation coarse and microsculpture very superficial to obliterate on discs.

4(5) Head red dorsally, except for a black median frontal carina and more or less infuscated genae. Antennae uniform pale red to dark brown but basal 1/2 scape pale. Genae barely shorter than eyes, GL/OL 0.82–0.94. Pronotum barely longer than wide, PL/PW 1.01–1.05. Body smaller, BL 19.4–20 mm. Apex of median lobe of aedeagus in lateral view as in couplet 2, yet short, very slightly rounded at tip and with a very acute dorsal tooth ( Fig. 26 ) ........... ........... 1a. G. japonica peregrina Dohrn, 1880 , stat.rest.

5(4) Body black but pale femora and sometimes also extreme base of antennal scape. Genae barely longer than eyes, GL/ OL 1.04–1.21. Pronotum longer, PL/PW 1.09–1.20. Body barely larger, BL 20–23.4 mm, macropterous to (rarely) apterous. Apex of median lobe of aedeagus in lateral view longer, subtriangular, with a minute, preapical, ventral tooth ( Fig. 28 ) .................... 2. G. batesi Andrewes, 1923

6(1) Body apterous and entirely black or with pale femora only. Pronotum mostly less than half wider at base than at apex. Elytra distinctly dilated apicad, with a slight yet more or less distinct impression spanning intervals 4–6 in basal third.

7(24) Elytra subglabrous due to all ridges with a single row of sparse and short setae on each side in at least apical two thirds. Legs black or with femora pale.

8(11) Femora pale. — Central and South Vientam.

9(10) Left paramere wide, triangular and rounded in apical third. Internal sac of aedeagus with a single, left, laterobasal bulb, and the body bent leftward and ventrad; 10–11 dorsomedial and 1–2 left lateral sclerites present. Genae slightly longer, GL/OL 1.27–1.43, pronotal base slightly narrower, PB/PA 1.27–1.45. — Quang Nam Province .... .................................................. 3. G. rufofemorata sp.n.

10(9) Left paramere narrower, trapezoidal. Internal sac of aedeagus with two laterobasal bulbs, left and right, and the body bent only leftward, i.e., set horizontally and perpendicular to median lobe; nine dorsomedial and 5–6 left lateral sclerites present. Genae shorter, GL/OL 1.17–1.32, pronotal base barely wider, PB/PA 1.32–1.50. — Gia Lai Province ............................... 5. G. dimorpha sp.n. (part)

11(8) Legs uniform black.

12(13) Body large, BL 26–28 mm. Head long, HL/HW 1.23–1.36, elliptic, with genae long, slightly rounded and converging much to neck. — Tam Dao, Vinh Phuc /Thai Nguen provinces ....... 10. G. tonkinensis Hovorka, 2019

13(12) Body smaller, BL<25.5 mm.

14(15) Elytra short and truncate combined apically (in male), EL/EW 1.64. Head moderately densely punctate in basal half. Left paramere subtriangular, with dorsal margin rounded and apex pointed. Internal sac of aedeagus without laterobasal bulbs, with six dorsomedial and two left lateral sclerites. — Lao Cai Province .............................. .......................................................... 9. G. truncata sp.n.

15(14) Elytra longer in male, mostly EL/EW 1.7–1.9; their apices obliquely truncate and thence slightly angulate combined (in both sexes). — Central Vietnam.

16(19) Genae shorter, GL/OL 1.17–1.32.

17(18) Pronotum barely narrower in male, PL/PW 1.11–1.20, otherwise as in couplet 8 … 5. G. dimorpha sp.n. (part.)

18(17) Pronotum barely wider in male, PL/PW 1.107. Left paramere very gently narrowed to and obliquely truncate at apex, with ventral margin straight and apex angulate. Internal sac of aedeagus with single, left, laterobasal bulb, six dorsomedial and two left lateral sclerites. — Kon Tum Province .................................. 4. G. konplongensis sp.n.

19(16) Genae longer, GL/OL 1.30–1.45.

20(21) Pronotum broadest closer to apex, PLw/PL 0.24–0.33. Left paramere elliptic, indistinctly desclerotized and truncate at apex. Many specimens have odd elytral ridges slightly yet distinctly stronger, wider and higher, than even ones. Internal sac of aedeagus with a conspicuous left laterobasal bulb, vestigial, short and wide, right one, nine dorsomedial and six left lateral sclerites. — Lam Dong Province ......................................... 6. G. variabilis sp.n.

21(20) Pronotum broadest more than a third from apex, PLw/ PL 0.35–0.39. Elytral ridges subequal.

22(23) Elytra short in female, EL/EW 1.59–1.61, and sparsely pubescent; head smaller, PW/HW 1.14–1.18. Impression on vertex conspicuous. — S-Vietnam (Dak Lak Province) ....................................................... 7. G. subglabra sp.n.

23(22) Elytra long in female, EL/EW 1.68–1.74, with pubescence fairly dense, more so in basal third, there forming 1–3 uneven rows on each side of longitudinal ridge. Head larger, PW / HW 1.23 – 1.27 . Vertex with no or superficial impression. Aedeagus with left paramere oblong, oviform, and subequally rounded at both dorsal and ventral margin; internal sac simple, with no laterobasal bulbs, eleven dorsomedial and 4–6 left lateral sclerites. — Central Vietnam (Ha Tinh and Vu Quang provinces) .............................................. 8. G.quadraticollis sp.n .

24(7) Elytra fasciated due to pubescence distinctly denser along even than along odd ridges. Legs black or nearly so. — North Vietnam.

25(26) Even elytral ridges less developed and much more densely pubescent than odd ones. Genae long, GL/OL 1.25–1.46, head wide, PW/HW 1.14–1.25. Elytra with distinct outer angles. Internal sac of aedeagus with three laterobasal bulbs, single left and two right, with eight medial and three or four left lateral sclerites; left paramere parallel-sided and obliquely subtruncate at apex. — Cao Bang Province ................................. 11. G. linearis sp.n.

26(25) Even and odd elytral ridges equally developed and less contrastingly pubescent. Genae shorter, GL/OL 1.08–1.21. Elytra with more or less rounded outer angles. Internal sac of aedeagus with single, right, laterobasal bulb, twelve medial and 7–8 left lateral sclerites.

27(28) Head slightly shorter and wider, HL/HW 1.13, PW/HW 1.20; pronotal base narrow, a third wider than apex. Left paramere triangular in apical half, with rounded apex; apex of median lobe with a ventromedial groove and fine carina at its bottom. — Nghe An Province ................................. ...................................................... 13. G. breviceps sp.n.

28(27) Head longer and slightly narrower, HL/HW 1.17–1.24, PW/HW 1.24–1.32; pronotal base wider, two fifths to half wider than apex. Left paramere barely narrower at a widely rounded apex than at base; apex of median lobe without ventromedial groove or carina. — Thanh Hoa Province.

..................................................... 12. G. sublineata sp.n.

Figures

Figs 22–31. Median lobe of aedeagus with adnate right paramere: 22–23 — Galerita orientalis from Assam, India; 24–25 — G. j. japonica from Japan; 26–27 — G. j.peregrina from Vu Quang NP; 28–29 — G. batesi from Karenni State, Myanmar; 30–31 — G. carinifrons from Bonthain, Celebes/ Sulawesi; 22, 24, 26, 28, 30 — left lateral aspect; 23, 25, 27, 29, 31 — right lateral aspect. Scale bar 1 mm. Рис. 22–31. СреднЯЯ долЯ Эдеагуса с приросшей правой парамерой: 22–23 — Galerita orientalis иЗ Ассама, ИндиЯ; 24–25 — G. j. japonica иЗ Японии; 26–27 — G. j.peregrina from from Vu Quang NP; 28–29 — G. batesi иЗ штата Карен, МьЯнма; 30–31 — G. carinifrons иЗ Бантаенг, Сулавеси; 22, 24, 26, 28, 30 — слева; 23, 25, 27, 29, 31 — справа. Масштаб 1 мм.