Platypalpus parvilamellatus Jonassen, 2021

Jonassen, Terje, Andersen, Trond, Salmela, Jukka, Suuronen, Anna & Vaesoja, Matti, 2021, Six new species of Empididae and Hybotidae (Diptera, Empidoidea) from Fennoscandia, Norwegian Journal of Entomology 68, pp. 174-186 : 182-184

Platypalpus parvilamellatus Jonassen sp. n.

( Figure 5A–C )

Type material. Holotype ♂, NORWAY: Finnmark, Porsanger, Stabbursdalen, Rørkulpen , N70.15215° E24.76686°, 28m .a.s.l. 17–26 July 2010, Malaise trap no. 5, Finnmarksprosjektet ( ZMBN) . Paratypes: 5♂♂, same data as holotype ; 2♂♂, same data as holotype except 26 July–25 August 2010 . 1♂, Finnmark, Alta, Gargiaelven at Gargia fjellstue, N69.80525° E23.48937°, 120m .a.s.l., 10–23 July 2010, Malaise trap no. 1, Finnmarksprosjektet ( ZMBN) ; 1♂, same data as previous except 23 July–07 August 2010 ; 1♂, same data as previous except 07–24 August 2010 . 3♂♂, Finnmark, Alta Gargiaveien, Gargiaelven at Storeng , N69.82277° E23.47884°, 90m .a.s.l., 10–23 July 2010, Malaise trap no. 2, Finnmarksprosjektet ( ZMBN) ; 6♂♂, same data as previous except 23 July–07 August 2010 ; 1♀, same data as previous except 07–24 August 2010 . 7♂♂, Finnmark, Kautokeino, Lahpoluoppal, Nahpoljohka , N69.21029° E23.76200°, 320m .a.s.l., 24 July–06 August 2010, Malaise trap no. 4, Finnmarksprosjektet ( ZMBN) . 6♂♂, Finnmark, Porsanger, Baukop, stream at Vuolit Gealbbotjavri , N70.20469° E24.90605°, 26m .a.s.l., 26 July–25 August 2010, Malaise trap no. 6, Finnmarksprosjektet ( ZMBN) . 9♂♂, Finnmark, Sør-Varanger, Sametijohka at Sameti , N69.40106° E29.71923°, 43m .a.s.l., 24 June–20 July 2010, Malaise trap no. 8, Finnmarksprosjektet ( ZMBN) . FINLAND: 1♂, Ks, Salla, Värriö , Kuntasjoki Ykj , N67.7494° E29.6169°, 29 June–29 July 2013, J. Salmela ( LMM: MH-EMPI-2013-0343) .

Additional material. NORWAY: Innlandet , Engerdal , Åsen , N61.885861 o E11.782833 o, 23 June–11 July 2016, 2♂♂; 11–21 July 2016, 1♂; 21 July–04 August 2016, 2♂♂, Malaise trap, Hedmarksprosjektet ( ZMBN). Innlandet , Rendalen , Sekserbua NE, N61.556056 o E11.168556 o, 23 June–11 July 2016, 1♂, Malaise trap, Hedmarksprosjektet ( ZMBN). Innlandet , Stor-Elvdal, Ottestad, N61.297111 o E11.277147 o, 20 July–06 August 2017, 1♂, light trap, Hedmarksprosjektet ( ZMBN). Trøndelag, Oppdal, Kongsvoll, 900m. a.s.l., 12 July 1992, 1♂, J. Skartveit (in coll. T. Jonassen). Trøndelag, Oppdal, Kongsvoll, Spraenbekken, 1020m.a.s.l., 01–10 August 1994, 1♂, J. Skartveit (in coll. T. Jonassen); 1♂, same data as previous except 950m. a.s.l . FINLAND: Ks , Salla , Värriö, Kuntasjoki, N67.7500° E29.6224°, 29 June–29 July 2013, 1♂, J. Salmela ( LMM: MH- EMPI-2013-0158); 1♂, same data as previous except N67.7494° E29.6169°, 04–29 June 2013, ( LMM: MH-EMPI-2013-0411). Obb, Kemijärvi, Pyhä-Luoston kansallipuisto, Huttuoja, N66.9983° E27.0265°, 01 July–08 August 2014, 1♂, J. Salmela ( LMM: MH-EMPI-2014-0609). Lkor, Savukoski, Törmäoja, N67.8357° E29.4540°, 11 June–09 July 2014, 1♂, J. Salmela ( LMM: MH- EMPI-2014-0662) .

DIffEREntIAL dIAgnOsIs. The species belongs in the Platypalpus ciliaris -group, and is most similar to P. confinis (Zetterstedt, 1842) . However, the two species can easily be separated as the male terminalia of P. parvilamellatus sp. n. is much smaller than in P. confinis . Particularly the left epandrial lamella is very small compared to the lamella in P. confinis . In addition, mid tibiae appear to be more curved in both sexes compared to P. confinis , with a larger apical spur. Further, P. parvilamellatus has a broader face than P. confinis , with the polished part of clypeus much broader and diamond-shaped, whereas this polished area is narrow and cigar-shaped in P. confinis .

Description. Male. Body length 2.2–2.6 mm, wing length 3.0– 3.4 mm.

Head. Black, greyish dusted. Frons about as broad as length of second antennal segment, widening above. Face about as wide as front of frons, dusted. Clypeus broadly polished, subequal in length to length of face. Polished part of clypeus almost diamond-shaped. Ocellar bristles minute, white. No discernible vertical bristles. Pubescence on head pale. Antennae black. Third antennal segment a little more than twice as long as broad, arista about twice as long as third segment. Palpus ovate, about as long as third antennal segment, yellow, covered with white pubescence, some apical hairs longer. Proboscis black.

Thorax. Black; mesonotum completely polished; humeri, scutellum and narrow lateral margins dusted. Pleura dusted, with whole of sternopleuron polished. Hairs and bristles pale.

Norwegian Journal of Entomology 68, 174–186 (2021)

Acrostichals biserial, inconspicuous, dorsocentrals uniserial, small, with the prescutellar pair larger, with only a few small hairs between dorsocentrals and margin of mesonotum. Notopleuron with 2–3 bristles, but these are difficult to distinguish from the subequal hairs. Humeri with 2–3 hairs longer than its general pubescence, without distinct humeral bristle. Scutellum with four bristles, the apical pair rather long, the other pair less than half as long.

Legs. All coxae, knees and basal half of hind femur yellow, remaining parts of the legs more or less brownish, tibiae darkest in apical ¾ of mid tibia and covered with an apical silvery pile. Tarsal segments becoming somewhat darker towards apex, last tarsal segment very dark brownish in all legs. Fore femur only indistinctly swollen, with two rows of fine ventral hairs. Mid femur moderately thickened, about as swollen as fore femur, with two rows of spine-like ventral bristles, posterior row strongest. No posteroventral bristles. Fore tibia cylindrical, thin, rod-like. Mid tibia ( Figure 5A ), distinctly curved, with small, darkened spur apically. Hind femur and tibia long and slender.

Wing. Faintly brownish with light brown veins. Crossveins contiguous. Vein closing anal cell slightly oblique, apical lower corner of anal cell <90º. Veins R 4+5 and M slightly curved, parallel in apical ¼. Halter pale, squama light brownish with pale hairs.

Abdomen. Black, shining, with pale pubescence. Hairs sparsely distributed, with terga a little more densely pubescent, hairs longest along posterior margin of tergum eight.

Terminalia ( Figure 5B–C ) small, in dried specimens often partly hidden within the abdomen. Lamellae small with left epandrial lamella notably minute. Cerci of normal size, consequently exceeding left epandrial lamella considerably. Right epandrial lamella with narrow, thumb-like process apically, encircled by some longer hairs. The small, left epandrial lamella with small triangular extension dorsally.

Female. Very similar to the male. Apical two abdominal segments and small cerci dusted.

Etymology. From Latin parvus, meaning small and lamellatus, with lamella, referring to the distinctive small lamella.

Distribution and habitat. The species is taken in northern and central Norway and in northern Finland. In northern Norway it was collected close to streams and rivers ( Ekrem et al. 2012).

Figures

FIGURE 5. Details of Platypalpus parvilamellatus Jonassen sp. n., male.A. Mid leg.B. Terminalia, dorsal view. C. Terminalia, ventral view.