Colias elis Strecker, 1885

Zhang, Jing, Cong, Qian, Shen, Jinhui, Opler, Paul A. & Grishin, Nick V., 2020, Genomic evidence suggests further changes of butterfly names, The Taxonomic Report of the International Lepidoptera Survey 8 (7), pp. 1-41 : 9-10

Colias elis Strecker, 1885 is a species distinct from Colias meadii W. H. Edwards, 1871

Tetracharis Grote, 1898 is a valid subgenus that includes Anthocharis cethura C. Felder & R. Felder, 1865 (Müller, 1764), Anthocharis midea (Hübner, [1809]) , and Anthocharis limonea (A. Butler, 1871)

The genomic tree of representative species of Anthocharis Boisduval, Rambur , [Duménil] & Graslin, [1833] (type species Papilio cardamines Linnaeus, 1758 ) including all type species of available genus-group names considered subgenera or junior subjective synonyms of Anthocharis revealed an unexpected but highly confident clade consisting of Anthocharis cethura C. & R. Felder, 1865 (type locality USA: California, Los Angeles Co. ), Anthocharis midea (Hübner, [1809]) (type locality USA:

Georgia, Wilmington Island ) and Anthocharis limonea (A. Butler, 1871) (type locality

Mexico) (Fig. 4 red). The clade is unexpected, because currently A. cethura is placed in the subgenus Anthocharis , while the Fig. 4. Subgenera Anthocharis (blue), Tetracharis (red)

other two species are placed in the subgenus and Paramidea (green) with Euchloe as outgroups. Paramidea Kuznetsov, 1929 (type species: Anthocharis scolymus Butler, 1866 ). Curiously, neither of these species in the red clade are monophyletic with the type species of subgenera they are currently assigned to: A. (Anthocharis) cardamines is in the blue clade (Fig. 4) and A. (Paramidea) scolymus is in the green clade. Therefore, the current classification is incorrect. Out of genus-group names that are available for the red clade, Tetracharis Grote, 1898 (type species Anthocharis cethura C. & R. Felder, 1865) is older than Falcapica Klots, 1930 (type species Papilio genutia Fabricius, 1793 , which is a junior homonym of Papilio genutia Cramer, 1779 , the oldest available name for this species is Mancipium midea Hübner, [1809] ). As a result, we treat Tetracharis as a valid subgenus, new status, that includes three species: A. cethura , A. midea , and A. limonea , making Falcapica its junior subjective synonym.