Physocephala caenoneura, Krober, 1939

Stuke, Jens-Hermann & Clements, David K., 2019, New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini, Bonn zoological Bulletin 68 (2), pp. 209-273 : 233-235

Key 4 – Identification of the Physocephala caenoneura species-group

1. Basal cell completely hyaline and almost completely lacking microtrichia, or with only very short microtrichia which are hardly visible and much less obvious than those on the remaining wing (e.g. Fig. 53 ); base of cell r 2+3 hyaline; hind femur at most with an indistinct dark spot ............................................. 2

– Basal cell at least with some dark colouration and sometimes completely black or brown, and almost completely covered with microtrichia similar to those on the remainder of the wing (e.g. Fig. 59 ); cell r 2+3 completely brownish infuscated (e.g. Fig. 59 ); hind femur may have a distinct blackish ring

234 Jens-Hermann Stuke & David K. Clements

which is sharply demarcated from the orange-brown colouration............................................................... 3

2. Tarsi orange-brown, not contrasting with tibiae; apical aristomere typically less than half length of projection on basal aristomere ( Fig. 48 ); antennae completely orange-brown ( Fig. 47 ); gena usually brownish, contrasting with yellow face ( Fig. 47 ); ♀ theca short in side view, distinctly shorter than wide ......................................... P. braunsi (Kröber, 1931)

– Tarsi dark brown, contrasting with brown tibiae ( Fig. 51 ); apical aristomere more than half as long as projection on basal aristomere ( Fig. 50 ); antennae black to brown, with a contrasting orange-brown scape (sometimes faded in old specimens) ( Fig. 50 ); gena usually yellow, like the face; ♀ theca long, almost as long as wide ( Fig. 54 )................................ .................................... P. caenoneura Kröber, 1939

3. Projection of basal aristomere elongated, about twice as long as apical aristomere ( Fig. 58 ); radial veins dark brown ( Fig. 59 ); basal cell only slightly darkened, obviously paler than cell r 2+3 ( Fig. 59 ) ...... .......................................... P. capensis Kröber, 1931

– Projection of basal aristomere not as elongated, not much longer than apical aristomere (e.g. Figs 44 , 61 , 65 ); radial veins sometimes light orange-brown; basal cell mostly as dark as cell r 2+3 (e.g. Fig. 45); base of cell r 4+5 and usually apex of basal cell hyaline, and therefore radial-medial crossvein within a hyaline patch (e.g. Fig. 60 )................................... 4

4. Characteristic dark mark at junction of discal medial-cubital crossvein and media ( Fig. 60 ); ♀ theca obviously short, with about 7 interrupted rows of black setae. ................. P. fenestralis ( Kröber, 1939)

– Wing colouration different, lacking dark mark at junction of discal medial-cubital crossvein and media, nor with hyaline spot surrounding radial-medial crossvein; ♀ theca longer, with more than 15 interrupted rows of black setae (e.g. Figs 43 , 64 ) ... 5

5. At least base of media and subcosta light yellowish-brown; scutum orange-brown, with three elongated blackish spots ( Fig. 63 ); katepisternum orange-brown; ♀ with black setulae on theca less dense, not obscuring underlying surface ( Fig. 64 ) ..................... ..................................... P. rufitarsis (Camras, 1962)

– Veins uniformly dark brown; scutum mainly blackish, with no distinct spots ( Fig. 42 ); katepisternum mainly black; ♀ with black setulae on theca denser, largely obscuring underlying surface ( Fig. 43 ) ..................... ................................. P. afenestralis ( Camras, 2001)

Figures

Figs 50–54. Physocephala caenoneura (Kröber, 1939) (♀, Nairobi). 50. Antenna, lateral view; 51. Leg with tarsi, dorsal view; 52. Frons, dorsal view; 53. Wing, dorsal view; 54. Theca, posterior view.

Figs 55–59. Physocephala capensis Kröber, 1931. 55. Theca, lateral view (♀, Zuurberg); 56. Theca, posterior view (♀, Zuurberg); 57. ♀ abdomen, dorsal view (♀, Zuurberg); 58. Arista, lateral view (♀, Zuurberg); 59. Wing, dorsal view (♀, Kwela lodge).

Figs 46–49. Physocephala braunsi Kröber, 2001 (♂, Marieps Mnt.). 46. Habitus, lateral view; 47. Head, lateral view; 48. Ari- stomeres, lateral view; 49. Wing, ventral view.

Figs 42–45. Physocephala afenestralis (Camras, 2001) (♀, Royal Natal National park). 42. Frons and scutum, anterodorsal view; 43. Theca, ventral view; 44. Arista, lateral view; 45. Wing, dorsal view.

Figs 60–62. Physocephala fenestralis (Kröber, 1939) (holotype). 60. Wing, dorsal view; 61. Arista, lateral view; 62. Wing, vena spuria in cell r4+5, lateral view.

Figs 63–65. Physocephala rufitarsis (Camras, 1962) (♀, Katberg). 63. Head and scutum, dorsal view; 64. Theca, posterior view; 65. Antenna, lateral view.