subsp. inodorus

Zhu, H., Roos, M. C. & Ridsdale, C. E., 2012, A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae), Blumea 57 (1), pp. 1-102 : 48-49

a. subsp. inodorus — Map 8

Lasianthus inodorus Blume (1826 –1827) 998; Miq. (1857) 322; (1869) 248; Merr. (1921) 577; Bakh.f. (1965) 339; H. Zhu (1998) 152; (2001a) 451; (2001b) 132; (2002) 77. — Mephitidia inodora (Blume) DC. (1830) 453. — Type: Blume s.n (holo L, barcode L0000690), Java.

Lasianthus tubiferus Hook.f.(1880) 183; Craib (1934) 219;Deb & M.Gangop. (1991) 305,f. 14; H. Zhu (1994) 59. — Type: Hooker & Thomson s.n. (lecto K; designated by Deb & Gangopadhyay 1991; isolecto A,L), India, Khasia.

Lasianthus poilanei Pit. (1924) 398. — Type: Poilane 234 (holo P; iso E, K), Mt Elephant , 1000 m, Cambodia.

Shrubs, 2 –3 m high; branches and branchlets glabrous or thinly puberulous. Leaves: blades elliptic to elliptic-lanceolate, 10– 20 by 2.5 –6 cm, coriaceous, glabrous above, glabrous or sparsely puberulous beneath, apex acuminate, base acuminate; nerves 6 –7 pairs, nervules parallel, nerves and nervules conspicuous beneath; petioles 6 –10 mm long, glabrous or sparsely puberulous. Stipules triangular or ovate-lanceolate, 3 – 5 mm long, glabrous, coriaceous. Cymes sessile; bracts persistent, 3 – 6 mm long, orbicular or ovate-orbicular, thickly coriaceous, glabrous or pubescent. Flowers sessile; calyx tubular, tube 2 mm long, glabrous, with 5 ovate-lanceolate lobes, puberulous outside; corolla 10 mm long, puberulous outside, villous inside. Drupes ovoid, 10 mm long, glabrous; pyrenes 5.

Distribution — India (northeast part), Bangladesh (eastern part), China (Yunnan), Vietnam, Cambodia, Thailand, Indonesia (Sumatra, Java).

Note — Subsp. inodorus has a wide, disjunct distribution. However, the specimens from mainland SE Asia and the ones from Sumatra and Java are almost identical so that clear variation within this subspecies cannot be recognized .