Lasianthus rigidus Miq.

Zhu, H., Roos, M. C. & Ridsdale, C. E., 2012, A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae), Blumea 57 (1), pp. 1-102 : 79-81

106. Lasianthus rigidus Miq. — Map 7

Lasianthus rigidus Miq.(1857) 321;Bakh.f.(1965)340;H. Zhu(2002) 75.—

Type: Junghuhn s.n. ( lecto L, designated by Zhu 2002,barcode L0000704 ), Java, Pengalengan . Paratypes: Unknown collector s.n. ( U, barcode U0006067 ) ; Junghuhn s.n. ( L, barcode L0305517 ), Java, Kananga .

Lasianthus tentaculatus Hook.f. (1880) 181; Deb & M. Gangop. (1991) 302, f. 12; H. Zhu (1994) 58; H.S. Lo (1999) 105. — Type: Wall. Cat. 8306 (holo K), India, Silhet .

Lasianthus humilis Elmer (1908) 519; Merr.(1923) 566. — Type: Elmer 9687 (holo lost; iso A, E, K, NY), Negros, Province of Negros Oriental, Cuernos Mountains , Dumaguete , Mar. 1908.

Shrubs, 1–1.5 m high; branchlets terete, robust, 2 – 3 mm diam, compressed and enlarged at nodes, glabrous or subglabrous. Leaves: blades obovate-oblong, obovate or oblong, 10–18 by 3.5 –7 cm, coriaceous, glabrous above, hirsute on midrib, nerves, and nervules beneath, apex shortly cuspidate-acuminate, base oblique, obtuse, slightly cordate or rounded, midrib and nerves flat above, prominent beneath, nerves 8– 9 pairs, ascending at an angle of 50–60°, curved to the margin; nervules distinct above, prominent beneath, parallel; petioles 3 –7 mm long, sparsely hirsute. Stipules persistent, triangular or subulate, 5 – 6 mm long, hirsute. Cymes sessile; bracts linear, linear-lanceolate to lanceolate, 9–12 mm long, numerous, hirsute. Flowers fascicular, sessile; calyx campanulate, 2– 3 mm long, sparsely hirsute outside, with 5 triangular 1 mm long teeth; corolla c. 10 mm long, puberulous outside, pubescent inside in the upper part. Drupes globose, glabrous, 4–5 mm diam, crowned by hirsute calyx teeth; pyrenes 5, smooth on the abaxial face.

Distribution — NE India, China (S Yunnan), Indonesia (Sumatra, Java), Philippines ( Negros Oriental).

Note — This species has leaves obovate-oblong or obovate, 12–18 cm long, 5 –7 cm broad, shortly cuspidate-acuminate at apex, the base oblique, obtuse, slightly cordate or rounded; stipules triangular or subulate, 5 – 6 mm long, hirsute; cymes sessile; bracts linear, linear-lanceolate to lanceolate, 9 –12 mm long, numerous, hirsute; drupes globose, glabrous with 5 pyrenes.

Figures

Map 7 Distribution of Lasianthus hirsutus (Roxb.) Merr.(¡) and L. rigidus Miq. (●).