Thyanta (Thyanta) rubicunda, Rider, 1991

Rider, D. A. & Chapin, J. B., 1991, Revision Of The Genus Thyanta Stal, 1862 (Heteroptera: Pentatomidae) I. South America, J. New York Entomol. Soc. 99, No. 1, pp. 1-77 : 11-16

Thyanta (Thyanta) rubicunda Rider, new species

Figs. 32-47, Map I

Description. Elongate ovate, dorsal surface pale to medium green or brown, often with pinkish-red markings between humeri, on apex of scutellum, and along lateral margins of pronotum, coria, and connexiva; punctures usually concolorous with surface.

Apex of head broadly rounded; outer jugal margins sinuous, not parallel, only slightly concave in front of eyes ( Fig. 33). Antennae pale brown to green, distal fourth of segment 3 dark brown, segments 4-5 entirely dark brown. Anterolateral pronotal margins in dorsal view concave, strongly denticulate for 2/J distance nearest head; humeral angles spinose, oriented anterolaterad, spines relatively short ( Fig. 32). Mesial angle of each pronotal cicatrice piceous. Coria densely and uniformly punctate; distal margins convex; costal angles angulate, reaching to middle of penultimate connexival segments ( Fig. 32); hemelytral membranes hyaline with a few scattered brown flecks. Connexiva narrowly exposed; posterolateral angle of each segment usually immaculate, sometimes minutely marked with black.

Ventral surface pale brown to green; punctures usually concolorous with surface. Rostrum pale brown to green, segment 4 mostly black, apex reaching between metacoxae. Femora and tibiae pale brown to green, tarsal segments and apex of each tibia darker. Postspiracular black spots lacking (except in brown form); posterolateral angles of each abdominal sternite at most minutely marked with black; anterolateral angles usually immaculate.

Mesial margins of basal plates in caudoventral view weakly concave, separated basally, almost contiguous apically; posterior margins sinuous, posteromesial angles broadly rounded ( Fig. 45). Distal end of sclerotized rod cone-shaped ( Fig. 46); spermathecal bulb digitiform, spermathecal duct forming distinct cylindrical structure below proximal flange ( Fig. 47). Pygophoral opening subtended by semicircular impression; posterior margin nearly straight with medial V-shaped emargination in caudal view ( Fig. 41); trisinuous in ventral and dorsal views ( Figs. 42, 43); posterolateral angles prominent in lateral view ( Fig. 44). Apex of each paramere spinose in ectal view ( Fig. 36); shaft rather robust at base with small tubercle ( Fig. 34); roughened, spiculate area on lateral surface linear ( Fig. 35). Each lateral conjunctiva! lobe of aedeagus with single diverticulum ( Fig. 37); dorsomedial lobe present ( Fig. 38).

Measurements. Total length 8.28-10. 72 (9.54); total width 5.83-7.02 (6.62); medial length ofpronotum 1.62-2.13 (l. 95). Medial length ofscutellum 3.48-4.53 (3.86); basal width 3.20-3.75 (3.42); width at distal end of frena 1.07-1.47 (1.40). Length ofhead l.72-1.90 (l.72); width 2.23-2.54 (2.30). Length of segments 1-5 ofantennae 0.40--0.44 (0.44), 0.92--0.98 (0.92), 0.92-1.03 (0.92), 1.03-1.10 (1.03), and 1.07-1.14 (1.14), respectively. Length of segments 2-4 of rostrum 1.32-1.47 (1.34), 0.74--0.88 (0.77), and 0.74--0.77 (0.74), respectively.

Holotype. 6 labeled (a) " Pocos, Antofagaste Prov. E. of Atacama Salt Lk. , Chile March 1955, Luis E. Pena, Collector" (b) "Thyantajuvenca Stal, Lutz '57." Deposited in the American Museum of Natural History (New York) .

Paratypes. 566, 1322. Labeled same as holotype except (b) "J C Lutz Collection 1961" (<;> USNM) ; (a) " Rayado Aconc . 18-VIII- 1960 " (b) " L. Campos colector" (6 IIAS) ; (a) " CHILE: San Pedro de Atacama, N. of Atacama Salt Lake , Antofagaster Prov. V- 1 -6-1964 " (b) "LE.Pena Collector" (2<;><;> AMNH) ; (a) " Estancia Castilla Vallenar " (b) " 8- Mayo 1969.-" (c) "}.Aranda Colector" (6 DAR;<;> IIAS) ; " Chile. Vallenar 3.VIl.86 En alfalfa Col. SAG " (6 IIAS); "CHILE Pudahuel Vegetaci6n 25.V.85 Col. R. Hevia" (6 IIAS) ; (a) " Chaca ( Chile?) 11 -5-55 L.E. Pena " (b) " Thyanta juvenca Stal, Det. J.C. Lutz" ( Q AMNH) ; (a) " Chile Mamina IX. 17.51" (b) "THOMAS F. HALSTEAD COLLECTION, California Academy of Sciences Accession" (2 CAS) ; (a) " Rio Lluta , Arica Dept. Tarapaca Prov., Chile Nov. 11-13, 1955; 500 Mt. Luis E. Pena , Collector" (b) "J C Lutz Collection 1961" (c) " Thyanta juvenca Stal" (2 USNM) ; (a) " LOMAS de PENUELAS LA SERENA -11- 1953" (b) " Thyanta chilensis H.S. Det. D.B.Thomas 1978" (2 MNHS); "LOMAS de PENUELAS. LA SERENA -11-1953" (2 MNHS) ; (a) " Los Andes , Chile " (b) " 17-V-79 Coll. G. Gordh" (2 UCR) ; (a) " Peiiueles 8-3-53 " (b) " Thyanta chilensis H.S. Det. D.B. Thomas 1978" (2 DBT) ; (a) " Arequipa Peru 10,28, '98" (b) "Herbert Osborn Collection" (<3 222 OSU), except 12 with (c) "may be patruelis St." and 12 with (c) "Arequipa Oct. 30,98." ( OSU) .

Distribution. Peru and Chile (Map l).

Comments. This species can be identified by the reduced amount ofblack markings on the abdominal venter, by the shape and orientation of each humeral spine, by the denticulation along the anterolateral pronotal margins, and usually by the pinkish coloration along the lateral margins of the body. The shape of the paramere is also unique within the nominate subgenus. The obtuse protuberance on the shaft of the paramere is reduced and nearer the base of the shaft.

Etymology. In Latin, rubicunda means pink-bordered, a character that many specimens of this species exhibit.