Pythonidina Grishin, 2019

Grishin, Nick V., 2019, Expanded phenotypic diagnoses for 24 recently named new taxa of Hesperiidae (Lepidoptera), The Taxonomic Report of the International Lepidoptera Survey 8 (1), pp. 1-16 : 6-7

Subtribe Pythonidina Grishin, 2019

Type genus: Pythonides Hübner, [1819] .

ZooBank registration:

CB890271-5483-4B5A-A7BC-27DBC5E23DE5

Diagnosis: The subtribe was defined as a clade from about 30 Mya that contains the genus Pythonides , and putatively synapomorphic DNA characters for this clade were given ( Li et al., 2019). Phenotypically, species in this subtribe would key to E.44a, E.49.1, or, if uncus undivided, then to E.37a or 40d in Evans (1953). A heterogeneous assembly of mostly small and frequently brightly patterned Hesperiidae not easily diagnosed morphologically. In all genera, palpi porrect, antennae not shorter than 1/2 costa, forewing without recurrent vein and hindwing costal margin not shorter than anal margin. Includes two subgroups by uncus morphology. If (1) uncus deeply divided ( Fig. 9 ), then 3rd segment of palpi as long as 2nd segment, antennal club nearly arcuate at its middle, apiculus not shorter than unbent portion of the club. If uncus undivided, and if (2) antennal apiculus shorter than unbent region of the club, then apiculus obtuse, not hooked, hindwing about triangular in shape, not quadrate, palpi not longer than head, or if (3) antennal club nearly arcuate at its middle and apiculus not shorter than unbent portion of the club, then 3rd segment of palpi shorter than 2nd segment.

Genera included: Ouleus Lindsey, 1925 , Zera Evans, 1953 , Quadrus Lindsey, 1925 , Gindanes Godman & Salvin, 1895 , Pythonides Hübner, [1819] , Haemactis Mabille, 1903 , Atarnes Godman & Salvin, 1897 , Eburuncus Grishin, 2012 , Milanion Godman & Salvin, 1895 , Paramimus Hübner , [1819], and Charidia Mabille, 1903 .

Parent Taxon: Tribe Achlyodini Burmeister, 1878 .

Figures

Fig. 9. Pythonidina. Long uncus arms in Eburuncus hierax (Hopffer, 1874), Peru (top left); palpi (long 3rd segment) of E. unifasciata (C. & R. Felder, 1867), Costa Rica (top right); antennae, bottom left: Ouleus calavius (Godman & Salvin, 1895), Panama (above), Pythonides lerina (Hewitson, 1868), French Guiana (below); palpi (short 3rd segment) of Gindanes brebisson panaetius Godman & Salvin, 1895, Panama (bottom right).